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The Alpine Geological History of the Hellenides, From the Triassic to the Present: Compression vs. Extension, the Dynamic Pair for the Orogen Structural Configuration. A Synthesis

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02 November 2023

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03 November 2023

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Abstract
Abstract: We present the Hellenic orogenic belt`s main geological structure and architecture of deformation, trying to reach a better interpretation of its geotectonic evolution during the Alpine orogeny. We based on our recent studies and experience about the deformational history of the Hellenides but also on the more modern views, published by other colleagues, concerning the Alpine geotectonic reconstruction of the Hellenides. The structural evolution started during the Permo-Triassic with the continental breaking of the Pangea super-continent and the birth of the Neotethyan ocean realm. Bimodal magmatism and A-type granitoid intrusions accompanied the initial stages of the continental rifting, followed by Triassic-Jurassic multiphase shallow- and deep-water sediment deposition on both formed continental margins. These margins were the Apulian margin, containing the Pelagonia, at the western part of the Neotethyan ocean and the European margin, containing continental parts of the Serbo-Macedonian and Rhodope massifs (upper Vertiskos-Kimi and middle Sidironero-Kerdylia units), at the eastern part of the Neotethyan ocean. Deformation and metamorphism are recorded in six main deformational stages from the Early-Middle Jurassic to the present day, getting started with an Early-Middle Jurassic Neotethyan intra-oceanic subduction and ensimatic island arc magmatism. Compression, nappe stacking, calc-alkaline magmatism and high-pressure metamorphic events related to subduction processes alternated successively through time with extension, orogenic collapse, medium- to high-temperature metamorphism, adakitic and calc-alkaline magmatism and partly migmatization, related to uplift and exhumation of deep crustal levels as tectonic windows or metamorphic core complexes. An S- to SW-ward migration of the dynamic peer compression vs. extension is recognized during the Tertiary Alpine orogenic stages in the Hellenides. In any case extension and crustal uplift follow compression and nappe stacking or compression and exten-sion act simultaneously at different parts of the orogen. We suggest that all ophiolite belts in the Hellenides originated from a single source and this was the Neotethyan Meliata/Maliac-Axios/Vardar ocean basin, parts of which obducted during the Mid-Late Jurassic on both continental margins, Apulian`s (containing Pelagonia) and European`s (containing units of Serbo-Macedonian/Rhodope nappes stack), W-SW-ward and E-NE-ward, respectively. In this case, the ophiolite nappes should be considered as far-traveled nappes on the Hellenides continental parts associated with the deposition of Middle-Late Jurassic ophiolitic mélanges in basins at the front of the adjacent ophiolite thrust sheets. The upper limit of the ophiolite emplacement is the Mid-Upper Jurassic, Callovian-Oxfordian, as shown by the deposition of the Kimmeridgian-Tithonian Upper Jurassic sedimentary carbonate series on the top of the obducted ophiolite nappes. The suture zone between the Pelagonian nappe and the External Hellenides as part of the Apulia plate has been dated about coeval in time with the Paleocene-Eocene to Oligocene SW-ward over thrusting (between 56 Ma and 32 Ma), along the Nestos thrust zone, of the Serbo-Macedonian/Rhodope Sidironero-Kerdylia unit on the tectonically lower-most Rhodope Pangaion unit. In this sense, the Nestos shear zone in the Rhodope province continues until the suturing between Pelagonian and External Hellenides and is the same with this. Therefore, the lower-most Rhodope Pangaion unit could be regarded, as the continuation of the marginal part of the Apulian plate (External Hellenides) that was underthrusted during the Paleocene-Eocene below the unified Sidironero-kerdylia unit and the Pelagonian nappe, following the Paleo-cene-Eocene subduction and closure of a small ocean basin at the western of the Pelagonian (Pindos-Cyclades ocean basin). Was preceded by the Late Cretaceous subduction of the Axios/Vardar ocean remnants below the European continental margin and the final closure of the Axios/Vardar ocean during the Paleocene-Eocene, which were associated with the overthrusting of the European origins Vertiskos-Kimi nappe on the Sidironero-Kerdylia nappe and subsequently the final collision of the European margin and Pelagonian fragment. In this scenario should be assumed, that the true Vardar/Axios suture zone should originate in the deep further to the East between the Rhodope units in the northern Rhodope province, while the Axios/Vardar zone in its today outcrops position should be considered as allochthonous. Subsequently, during synorogenic Oligocene-Miocene extension, the lower-most Rhodope Pangaion unit exhumed as a metamorphic core complex below the Serbo-Macedonian/Rhodope nappes, which were successively exhumed, from the higher to the middle, during Paleocene-Eocene and Eocene-Oligocene extension, respectively. A retreating subduction zone associated with rollback of the subducted lithosphere under the Pelagonian and the other Internal Hellenides nappes stack and/or slap break off and mantel delamination could well explain the Tertiary synorogenic extensional tectonics in the Internal Hellenides, taking place simultaneously with compression in the External Hellenides and the Hellenic foreland or accretionary wedge.
Keywords: 
Subject: Environmental and Earth Sciences  -   Geophysics and Geology

1. Introduction

Today, the Hellenides shows an arcuate orogenic belt extending from Dinarides in the northern to Taurides and Menderes and Sakarya massifs in the eastern, traditionally subdivided into the External and Internal Hellenides zones. (Figures 1, 2). They form a branch of the broader Alpine orogenic belt in Eurasia, resulting from the convergence and final continental collision of the European and Apulian plates in a complicated, multiphase Alpine deformational regime, where subduction processes, compression and nappes stacking alternated successively with extension, tectonic denudation and crustal exhumation from the Permo-Triassic to the recent. In any case, in this tectonic framework, some questions about the structural and geotectonic evolution of the Hellenides remain under debate until today, or several controversial interpretations have been proposed for them. E.g., how many Mesozoic Tethys ocean basins, one or more, existed between the European and Apulian continents and what were the origin and the obduction direction of the ophiolite nappes on the continental segments (e.g., Meliata/Maliac-Axios/Vardar ocean, Pindos/Mirdita ocean, etc.). Furthermore, the paleogeographic and tectonic origin of the old Paleozoic, pre-Alpine crystalline Pelagonian and Serbo-Macedonian/Rhodope continental segments with their Triassic-Jurassic sedimentary cover incorporated in the Hellenides are also open questions (e.g., a critical element on these questions is the geotectonic and paleo-geographic position of the tectonically lowermost Rhodope Pangaion unit and the other geological units constituting the Serbo-Macedonian/Rhodope nappes stack)
Several geotectonic models with controversial conclusions have been suggested until today to explain the geotectonic evolution and paleogeographic conditions of the Hellenides (e.g., [1,2,3,4,5,6,7,8,9,10,11,12,13,14,15,16,17,18,19,20,21,22,23,24]) and references therein). These controversial views are presented in detail below in the descriptions of the isopic geological units or zones and the structural evolution of the Hellenides.
The goal of this work was to present one aspect of the structural architecture and geodynamic evolution of the Hellenides, as well as the oceanic source and the emplacement direction of the obducted ophiolites and finally, the paleogeographic origin of the pre-Alpine continental blocks incorporated in the Hellenic orogenic belt. We based our conclusions not only on our recent studies and experience about the deformational history of the Hellenides but also on the more modern views, published by other colleagues, concerning the Alpine geotectonic reconstruction of the Hellenides.
Figure 1. The main structural domains of Dinarides and Hellenides as parts of the broader Alpine orogenic belt in Europe. Insert: The Alpine orogenic belt in Europe (modified after [7]).
Figure 1. The main structural domains of Dinarides and Hellenides as parts of the broader Alpine orogenic belt in Europe. Insert: The Alpine orogenic belt in Europe (modified after [7]).
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Figure 2. Geological map of the Hellenides with the main tectonic zones and their possible continuation to the East in Turkey (modified after [7,9,12]).
Figure 2. Geological map of the Hellenides with the main tectonic zones and their possible continuation to the East in Turkey (modified after [7,9,12]).
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2. Geological Setting

Maintaining the old subdivision of the Hellenides in isopic zones (i.e., [25,26,27]), the External and Internal Hellenides are composed, from the West to the East, by the following main tectonostratigraphic domains:
2.1. External Hellenides: I. Paxos zone, II. Ionian zone, III. Gavrovo zone and IV. Pindos zone, possibly including the Koziakas unit.
2.2. Internal Hellenides: I. Pelagonian zone or nappe and ?Parnassos zone II. ?Sub-Pelagonian zone, III. Axios/Vardar zone, IV. Cycladic massif, V. Circum-Rhodope belt, VI. Serbo-Macedonian massif, VII. Rhodope massif.
The External Hellenides are mainly built up by Mesozoic and Cenozoic deep-sea and shallow-water sedimentary rocks, characterized by continuous sedimentation processes terminated in Paleocene to Miocene flysch deposition. They form a complicated SW- to SSW-verging, thin-skinned thrust and fold belt of the Paleogene to Neogene age without any important metamorphism (Figures 3, 4; [25,26,28,29,30] and references therein). Due to Oligocene-Miocene internal underthrusting between litho-stratigraphic domains of the External Hellenides, a high-pressure/low-temperature metamorphic belt (HP/LT) was developed, which is today exhumed in the southern Peloponnese and Crete island. It is recorded in the Plattenkalk, Phyllite-Quartzite and Tripali units (Figures 1, 2, 3, 4, 5; [31,32,33,34,35]).
On the other hand, the Internal Hellenides are composed of Paleozoic and older basement rocks covered by Triassic-Jurassic carbonate platform sediments, as well as outer shelf and shelf edge sedimentary series, on which the Neo-Tethyan ophiolites realm were initially obducted during the Mid-Late Jurassic. Internal Hellenides are characterized by a multiphase tectonic history and metamorphism during the Alpine orogeny, from the Jurassic to the Tertiary (D1 to D6 events; they are in detail described in the chapter “Architecture of deformation and structural evolution). The Internal Hellenides overthrust the External Hellenides during the Eocene-Oligocene (Figures 3, 4, 5; [6,18,19,21,30,36,37,38,39,40,41,42,43] and references therein).
A Paleocene-Eocene high-pressure belt characterizes the tectonic contact between the External and Internal Hellenides zones exhumed in the Olympos-Ossa and Cyclades provinces directly under the Internal Hellenides Pelagonian nappe (Figures 1, 2, 3, 4; [38,39,44,45,46,47,48,49]). Additionally, high- to ultrahigh-pressure metamorphic conditions of Jurassic-Cretaceous and Paleocene-Eocene age are also described for the Serbo-Macedonian/Rhodope metamorphic province of the Internal Hellenides [50,51,52,53,54,55,56,57]. Recent works recognize a Late Cretaceous high-pressure belt in the Rhodope province [58,59]. Late orogenic Eocene to Early Miocene molassic-type (turbidites) basins, such as the Mesohellenic trough (Figures 1, 2, 3, 4; [60,61,62,63,64,65]) and the Thrace basin (Figures 1, 2, 3, 4; [7,30,66,67]) were developed locally on the top of the External and Internal Hellenides structural sequences. Finally, post-orogenic Neogene-Quaternary intramontagne and other terrestrial sedimentary basins cover unconformably in many places, all the pre-Alpine and Alpine Helenides tectonostratigraphic domains (Figures 1, 2, 3, 4; e.g., [68,69,70,71,72,73]).
Figure 3. Geological map of Northern Greece compiled after [5–7,9–13,15–21,24–27,30,34,36,38,39,40,43,48,67,68 and references therein]. Qu=Quaternary, Ne=Neogene, Mi=Miocene, Ol=Oligocene, Eoc=Eocene, Pal=Palecene, Cr=Cretaceous, Jr=Jurassic, Tr=Triassic, Pm=Perm, Pa=Paleozoic and older, L=Late, E=Early. A-A` cross-sections in Figure 4.
Figure 3. Geological map of Northern Greece compiled after [5–7,9–13,15–21,24–27,30,34,36,38,39,40,43,48,67,68 and references therein]. Qu=Quaternary, Ne=Neogene, Mi=Miocene, Ol=Oligocene, Eoc=Eocene, Pal=Palecene, Cr=Cretaceous, Jr=Jurassic, Tr=Triassic, Pm=Perm, Pa=Paleozoic and older, L=Late, E=Early. A-A` cross-sections in Figure 4.
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2.1. External Hellenides

2.1.1. Paxon Zone

The Paxon zone forms the most external zone of the External Hellenides and continues to the North in the Albanides and Dinarides (Figures 1, 2, 4). It is characterized by continuous, neritic carbonate sedimentation from the Mid-Upper Triassic to the Oligocene-Miocene without typical flysch deposition, which is exposed in all other external Hellenides zones. Thin strata of siliceous sediments and shales are locally interbedded with the Jurassic carbonate succession. Furthermore, evaporites are the older, lower-most litho-stratigraphic sequence of the zone [26,74,75,76,77,78,79].

2.1.2. Ionian Zone

The Ionian zone (Figures 1, 2, 3, 4) is built up by a continuous sedimentary series from the Triassic to the Oligocene-Miocene. It is also recognized to the North in the Albanides and Dinarides orogenic belts. Until the Early-Middle Liassic, the sedimentation was characterized by neritic calcareous deposits, terminating with the characteristic Pantokrator white limestone series of the Liassic age. On the contrary, from the Middle Jurassic until the Eocene, the sedimentary conditions changed to pelagic with radiolarites and pelagic limestones deposition and finally ended with an Oligocene-Miocene flysch deposition. As in the Paxon zone, evaporites are also the older, lower-most litho-stratigraphic sequence of the Ionian zone [25,26,30].
Equivalent to the Ionian sediments seems to be the Plattenkalk unit in the Peloponnese and the Crete island, cropping out tectonically together with the Jurassic age`s Tripali unit under the Phyllite-Quartzite unit and the Gavrovo zone (Figures 2, 4, 6; [32,33,34,80,81,82]). The Plattenkalk, the Tripali and the Phyllite-Quartzite units have been affected by the Oligocene-Miocene HP/LT metamorphic event of the External Hellenides, while the overlain Gavrovo zone remains unmetamorphosed here (Figures 2, 6). The Plattenkalk are rich in metamorphic aragonite in the marbles and Fe-Mg-carpholite in the intercalated thin metabauxites layers. Additionally, in the Phyllite-Quartzite unit, it has been recognized that glaucophane is also associated with Fe-Mg-carpholite. The Plattenkalk unit, together with the Tripali and Phyllite-Quartzite units, were uplifted and exhumed rapidly under Early-Middle Miocene extensional tectonics and isothermal decompression (Figure 6; [3,4,5,6,7,8,9,10,11,12,13,14,15,16,17,18,19,20,21,22,23,24,25,26,27,28,29,30,31,32,33,34,35,82,83,84]).
The Ionian zone is thrust over the Paxos zone towards the West during the Mid-Late Miocene (Figures 2, 4).

2.1.3. Gavrovo Zone

The Gavrovo zone is characterized by continuous Triassic to Eocene neritic, platform sedimentation on the Apulia continental passive shelf, terminated by the deposition of an Eocene-Oligocene flysch (Figures 1, 2, 4, 5; [25,26,30,36,85]). It is also exhumed to the North in the Albanides and Dinarides. The western parts of the Gavrovo zone in the External Hellenides appear unmetamorphosed and intensively imbricated during the Oligocene-Miocene. However, its more eastern parts that crop out in the Internal Hellenides as tectonic windows or metamorphic core complexes in Olympos-Ossa, Rizomata, Almyropotamos-Attica and Cyclades provinces (Figures 2, 3, 4, 7), beneath the Pelagonian nappe and the glaucophane-bearing Paleocene-Eocene high-pressure blue-schists belt (Ampelakia and Cyclades units) show evidence of a Tertiary (Oligocene-Miocene) low- to high-grade metamorphism related to extension and exhumation. So, e.g., in Olympos-Ossa province, only low-grade metamorphism is recorded without any significant reheating (Figure 8a; [5,38,39,47,48,86,87]). On the contrary, in Almyropotamos-Attica and Cyclades provinces, significant reheating, high-temperature metamorphism, migmatization and abundant granitoid intrusions occurred during the Oligocene-Miocene (Figure 8b; [45,88,89,90,91,92,93,94,95]).
Figure 4. a. and b. Representative geological cross-sections through northern Greece, showing the geometry of deformation and structural architecture of the Hellenic orogenic belt as it was created by the Alpine orogenic processes from the Jurassic to the recent (the deformational events are in detail described in the chapter “Architecture of deformation and structural evolution”). The tectonic windows of Olympos-Ossa and Rhodope Pangaion are shown. MHT = Mesohellenic trough. Legend and abbreviations as in Figure 3 (modified after [6,7,17,24,48,57,67 and references therein]. 26-15 Ma=cooling/exhumation data, 43 Ma AFT=apatite fission- track ages of the several Serbo-Macedonian/Rhodope tectonic nappes [225].
Figure 4. a. and b. Representative geological cross-sections through northern Greece, showing the geometry of deformation and structural architecture of the Hellenic orogenic belt as it was created by the Alpine orogenic processes from the Jurassic to the recent (the deformational events are in detail described in the chapter “Architecture of deformation and structural evolution”). The tectonic windows of Olympos-Ossa and Rhodope Pangaion are shown. MHT = Mesohellenic trough. Legend and abbreviations as in Figure 3 (modified after [6,7,17,24,48,57,67 and references therein]. 26-15 Ma=cooling/exhumation data, 43 Ma AFT=apatite fission- track ages of the several Serbo-Macedonian/Rhodope tectonic nappes [225].
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Figure 5. Schematically, the tectonostratigraphic column of the Hellenides. The several Alpine deformational events and their actions are shown (they are in detail described in the chapter “Architecture of deformation and structural evolution”). Abbreviations as in Figure 3.
Figure 5. Schematically, the tectonostratigraphic column of the Hellenides. The several Alpine deformational events and their actions are shown (they are in detail described in the chapter “Architecture of deformation and structural evolution”). Abbreviations as in Figure 3.
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The pre-Alpine basement of the Gavrovo carbonate platform (Apulia plate) is exhumed as metamorphic core complexes in the Cyclades province beneath the Gavrovo carbonate series, the Cyclades blue schists unit and the overlain remnants of the Pelagonian nappe with the obducted Neotethyan ophiolites (Figures 2, 4; [9,89,93,96,97,98,99,100]). It is possibly continued further to the East in western Turkey in the Menderes metamorphic core complex as its deeper structural level (Figure 2; [9,98]). Both the Cyclades and Menderes provinces show at least an equivalent tectonostratigraphic setting and Tertiary tectonic history. From the bottom to the top are recognized: a pre-Alpine basement, covered by neritic limestones or marbles in places of Triassic to Eocene age (and they are the Gavrovo carbonate platform), overthrust by the Paleocene-Eocene Ampelakia-Cyclades high-pressure belt, and topped tectonically by the Pelagonian nappe or the Lycian nappe in the Hellenides and the Menderes area, respectively. Triassic granitoid intrusions in both provinces are another common point among these. Oligocene-Miocene granitoid intrusions also occur in both the Menderes and Cyclades domains, but in different amounts. They are more in the Cyclades massif (Figure 2; [9,10,15,45,89,95,99,101,102,103,104,105,106,107,108,109]). Furthermore, kinematic indicators show real similarities in both areas of the Cyclades and Menderes. In the Cyclades, the Oligocene-Miocene extension direction evolves mainly N- to NE-ward and is related to exhumation processes (Figure 9), as it is also described for the Menderes region. Additionally, S- to SW-ward sense of movements have also been described for both areas (Figure 9; [9,11,89,93,98,101,102,103,110]).
Nevertheless, some lithological and structural differences should also be referred to here between the two regions, leaving some question marks on the correlation of the two areas and raising the need for further detailed research for the correct answer. For example, the absence of the Pelagonian nappe in Menderes province, where the Lycian nappes may be the equivalent structural sequence of the Pelagonian nappe, taking the same tectonic position with it [9,10,99].
Moreover, the Menderes basement forms a Pan-African continental segment [98,101,102,105,110]. One more difference is the obduction age of the ophiolites. In the Hellenides, it is of Mid-Late Jurassic age and in the Menderes province, it is of Late Cretaceous age ([80,112,113] and references therein). Another difference is the larger amount of basement rocks in the Menderes massif than in the Cycladic massif (Figure 2; [45,90,99,101,102,105]).
Recent works by [17,24,114] support the continuation of the Gavrovo carbonate platform as part of the Apulian`s plate passive margin until the Rhodope metamorphic province, named the Pangaion metamorphic core complex. It is exhumed as a tectonic window below the metamorphic Serbo-Macedonian/Rhodope tectonic nappes or terranes of the Internal Hellenides (Figure 2).
The Gavrovo zone is thrust over the Ionian zone towards the West during the Oligocene-Miocene (Figures 1, 2, 4).

2.1.3. Pindos Zone

The Pindos zone is continued to the North in the Albanides and Dinarides orogenic belts, while to the South, it is recognized as small, residual tectonic nappes in Crete island, as well as in some Aegean islands East of Crete (Figures 1, 2, 3). The Pindos zone is characterized by deep-sea sediments (such as pelitic, siliceous and pelagic carbonate deposits) of Triassic to late Late Cretaceous age, overlain by a Paleocene-Eocene flysch which shares in places the typical features of wild flysch. Nevertheless, the Pindos first depositional materials are clastic neritic sediments of Early-Middle Triassic age, with volcanoclastic products intercalated. An occurrence of Early Cretaceous flysch-type deposits (Pindos “first flysch”) in between the deep water Pindos strata remains until today under debate. They are composed of alternations of thin layers of red marls and radiolarian cherts, marly limestones, pelites and green fine- to coarse-grained sandstones with ophiolitic material, as well as ophiolite pebbles [25,26,28,29,30].
The geotectonic-paleogeographic position of the Pindos zone is widely controversial. It is believed to have developed at the western Pelagonian margin, either as an ocean basin opened progressively due to the Permo-Triassic continental rifting of Pangea [3,14,19,25,113,115,116] or as a deep sea basin formed on a thinned continental crust [113]. Moreover, recent works by [17,23,24,59,117] suggest the opening of a narrow ocean basin during the Late Cretaceous, named the Pindos-Cyclades ocean, which divided the Pelagonian of Apulia at this time. This small ocean, as part of the entire Pindos zone, was subducted NE-ward during the Paleocene-Eocene under the Pelagonian fragment and the Internal Hellenides nappe stack, where it metamorphosed under high- to ultra-high-pressure metamorphic conditions and then overthrusted during the Eocene-Oligocene on the Apulian margin together with the Pelagonia [5,36,38,39,47,48,86,87].
In any case, the entire Pindos zone overthrusts to W- to SW-ward the Gavrovo carbonate platform during the Eocene-Oligocene. It shows a similar structural position with the subducted under the Pelagonia Paleocene-Eocene blue-schist belt that is exhumed in the Olympos-Ossa and Cyclades provinces tectonically direct above the Gavrovo zone.
Figure 6. a. P/T/t tectono-metamorphic path and exhumation history of the Oligocene-Miocene high-pressure belt (Plattenkalk, Tripali and Phyllite-Quartzite units) in Crete island [32,33,296,297]). b. Field- and macroscale photos of the geological units and deformational structures on the Crete island (I to IX). The sense of shear is shown by arrows and it is related to the Early-Middle Miocene extension and exhumation of the Oligocene-Miocene HP/LT metamorphic belt of the External Hellenides (Peloponnese and Crete).
Figure 6. a. P/T/t tectono-metamorphic path and exhumation history of the Oligocene-Miocene high-pressure belt (Plattenkalk, Tripali and Phyllite-Quartzite units) in Crete island [32,33,296,297]). b. Field- and macroscale photos of the geological units and deformational structures on the Crete island (I to IX). The sense of shear is shown by arrows and it is related to the Early-Middle Miocene extension and exhumation of the Oligocene-Miocene HP/LT metamorphic belt of the External Hellenides (Peloponnese and Crete).
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Therefore, the Pindos zone in the External Hellenides possibly represents the non-metamorphic part of the blue-schists that has escaped the subduction under the Pelagonian continental fragment and continues until this Paleocene-Eocene high pressure belt (Figures 1, 2, 3, 4; [5,6,23,38,39,47,48,80,118,119,120]). On the contrary, [121] regard this blue-schist belt as of Pelagonian origin.
The Koziakas Unit in central Greece (Figures 2, 3) is composed from the bottom to the top of pelagic carbonate sediments intercalated with radiolarian cherts and shales of Middle Triassic-Early Jurassic age, Jurassic thick-bedded oolithic limestones and redeposited mass–flow sediments of Early Cretaceous age and also Early Cretaceous flysch-type sediments equivalent to the Beotian flysch and the Pindos “first flysch”. On the Beotian flysch are to W-SW-ward overthrust the Triassic and Jurassic sequences of the Koziakas unit, forming additionally the whole Koziakas unit an overturned W-ward vergent mega-fold. Moreover, in the stratigraphic column of the Koziakas unit are also described shallow-water Late Cretaceous limestones and micro-breccia deposits (Thymiana limestones) and finally, Paleocene-Eocene flysch. According to many authors, the Koziakas unit is a sedimentary series with continued sedimentation from the Triassic to the Eocene equivalent to the Pindos zone (Figures 2, 3; [19,25,122,123,124,125,126,127]). Nevertheless, on the Triassic-Liassic deep-see sedimentary series of the Koziakas unit are tectonically emplaced Middle-Late Jurassic ophiolitic mélanges and the Neotethyan ophiolites. It raises some questions about the existence of continued sedimentation in the unit as well as its geotectonic position. Furthermore, according to other authors, the Koziakas unit has been regarded as an individual tectonostratigraphic unit of unknown origin, thrust during the Paleocene-Eocene on the Pindos unit from the East to the West ([128] and references therein), so that its geotectonic setting, as well as paleo-geographic position, remain under discussion.
Figure 7. Geological-structural map and representative cross-section of the tectonic boundary between the Pelagonian nappe and the Axios/Vardar zone at Vermion and Pieria Mts. The Rizomata window, equivalent to the Olympos-Ossa window, as well as the Late Jurassic tectonically duplication of the Pelagonian nappe and the Paleocene-Eocene Almopian tectonic sheets, are shown (based on [43,131,154]). Abbreviations as in Figure 3.
Figure 7. Geological-structural map and representative cross-section of the tectonic boundary between the Pelagonian nappe and the Axios/Vardar zone at Vermion and Pieria Mts. The Rizomata window, equivalent to the Olympos-Ossa window, as well as the Late Jurassic tectonically duplication of the Pelagonian nappe and the Paleocene-Eocene Almopian tectonic sheets, are shown (based on [43,131,154]). Abbreviations as in Figure 3.
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Figure 8. a. and b. P/T/t tectono-metamorphic path and exhumation history of the Paleocene-Eocene high-pressure belt (blue schists), (a) in Olympos-Ossa and (b) Cyclades areas, respectively [45,46,47,48,88]); c. Field- and microscale photos of the geological units and deformational structures, (I to V) in the Olympos-Ossa and (VI to VIII) in Cyclades provinces: I and II. The Olympos normal detachment zone, along which detached SW-wards the Pelagonian nappe pile with the Late Jurassic obducted ophiolites and the overlain Late Jurassic-Cretaceous sedimentary series, resulting in the uplift and final exhumation of the high-pressure blue schists and the Olympos carbonate unit (Gavrovo zone); III, IV and V. Meso- and microscale features of deformation of the Ampelakia high-pressure belt. S-C fabrics and shear bands in the blueschists (III and IV), pyroxene σ-clasts in the intercalated metabasites of the blueschist unit, glaucophane growing in the pressure shadows of the pyroxenes during the high-pressure process (V), X-Z sections, the main sense of shear top-to-SW during the exhumation processes; VI and VII. Fold structures in the Cycladic metamorphic belt; VIII. Megalodonts bearing recrystallized limestones of the Triassic-Jurassic Emporio carbonate unit in Santorini island [103]).
Figure 8. a. and b. P/T/t tectono-metamorphic path and exhumation history of the Paleocene-Eocene high-pressure belt (blue schists), (a) in Olympos-Ossa and (b) Cyclades areas, respectively [45,46,47,48,88]); c. Field- and microscale photos of the geological units and deformational structures, (I to V) in the Olympos-Ossa and (VI to VIII) in Cyclades provinces: I and II. The Olympos normal detachment zone, along which detached SW-wards the Pelagonian nappe pile with the Late Jurassic obducted ophiolites and the overlain Late Jurassic-Cretaceous sedimentary series, resulting in the uplift and final exhumation of the high-pressure blue schists and the Olympos carbonate unit (Gavrovo zone); III, IV and V. Meso- and microscale features of deformation of the Ampelakia high-pressure belt. S-C fabrics and shear bands in the blueschists (III and IV), pyroxene σ-clasts in the intercalated metabasites of the blueschist unit, glaucophane growing in the pressure shadows of the pyroxenes during the high-pressure process (V), X-Z sections, the main sense of shear top-to-SW during the exhumation processes; VI and VII. Fold structures in the Cycladic metamorphic belt; VIII. Megalodonts bearing recrystallized limestones of the Triassic-Jurassic Emporio carbonate unit in Santorini island [103]).
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Figure 9. Geometry of the kinematics of the ductile deformation (sense of shear) during the Tertiary extensional tectonics in the Hellenides that progressively migrated towards the SW (modified after [5]).
Figure 9. Geometry of the kinematics of the ductile deformation (sense of shear) during the Tertiary extensional tectonics in the Hellenides that progressively migrated towards the SW (modified after [5]).
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2.2. Internal Hellenides

2.2.1. Pelagonian Zone or Nappe

The Pelagonian nappe or Pelagonian zone, is composed from the top to the bottom of: I. a Triassic-Jurassic carbonate platform sequence; II. a volcano-sedimentary Permo-Triassic series characterized by bimodal magmatism: and III. a Paleozoic or older pre-Alpine crystalline basement, composed of gneisses, amphibolites and schists, intruded by Carboniferous calc-alkaline (~300 Ma) and Triassic A-Type (~240 Ma) granitoid. Mesozoic or Cenozoic magmatic activity has not been recorded in the Pelagonian basement (Figures 3, 4, 5; [3,6,129,130,131,132,133,134]). The Pelagonian nappe was divided by [6], due to tectonic internal thrusting during the Late Jurassic into a tectonic lower and a tectonic upper Pelagonian segment, both showing the same tectonostratigraphy column as described just above (Figures 3, 4, 5). Obducted Neo-Tethyan ophiolites, imbricated with Mid-Late Jurassic ophiolitic mélanges, occur in many places on the Triassic-Jurassic Pelagonian carbonate cover or seco-ndary, directly on the pre-Alpine basement (Figures 1, 2, 3, 4). The ophiolites obduction took place during the Mid-Late Jurassic, following the intra-oceanic subduction/-s in the Neo-Tethyan ocean basin/-s [6,13,19,21,135,136,137]. The Pelagonian nappe, together with the obducted ophiolite belt, crop out mainly in continental Greece, overlying tectonically the Gavrovo zone and the Paleocene-Eocene Ampelakia-Cyclades high-pressure belt, which are exhumed as tectonic windows below the Pelagonia, as in detail described above. The Pelagonian nape is also recognized on top of the Cyclades metamorphic complex as small, far-traveled relics and on Crete island as the Asterousia nappe (Figures 2, 6; [23,32,33,76,80,118,138]). Furthermore, clasts of Pelagonian nappe origins are also found in Early Miocene conglomerates deposited above detachment faults in the Cyclades province, showing really that the Pelagonian nappe once covered a large part of the Aegean region and finally eroded or tectonically denudated [95,139,140,141]. The basement gneissic rocks and their Triassic-Jurassic carbonate cover of the Paikon subzone in the Axios/Vardar zone at Tzena and Paikon Mts were regarded by [142] as a tectonic window of Pelagonian origin. Nevertheless, the Pelagonian nappe is also recognized to the North in the Dinarides, while its continuation to the East in Turkey remains under discussion today. As referred to above in the description of the Gavrovo zone, parts of the Lycian nappes in southwestern Turkey maybe, could form the relics of the Pelagonian nappe further to the East (Figure 1; [3,6,9,113]).
Furthermore, Late Jurassic strongly eroded carbonate platform sediments, Early Cretaceous mass flows and flysch-like deposits, as well as transgressive Late Cretaceous shallow-water carbonate sedimentary series, rest in places on top of the obducted Neo-Tethyan ophiolite realm or directly on the exhumed Pelagonian Triassic-Jurassic platform carbonate. (Figures 5, 10; [13,21,27,143,144,145,146,147,148,149,150]).
A detailed P/T/t-path of the complicated, polyphase Alpine tectono-metamorphic history of the Pelagonian basement rocks, which is in detail discussed in numerous works by [13,21,27,143,144,145,146,147,148,149,150], is displayed in Figure 11a, b, c. It shows: I. High-pressure conditions during the Middle-Late Jurassic, associated with thrusting and crustal thickening, II. Amphibolite to greenschist facies retrogression during the Late Jurassic-Early Cretaceous and extensional crustal uplift, III. Late Early Cretaceous (Barremian-Aptian) greenschist facies prograde metamorphism related to compression and SW-ward thrusting, IV. Late Cretaceous cooling/exhumation for the structurally upper Pelagonian parts and brittle compressional deformation and thrusting during the Paleocene-Eocene. It was followed from the Oligocene-Miocene to the recent by extension. In contrast, for the structurally lowermost Pelagonian parts, near their tectonic contact with the underlain Paleocene-Eocene blue schists belt or the External Hellenides (Gavrovo zone), ductile deformation under high-pressure conditions is followed by low-grade metamorphic retrogression under extension and strongly mylonitization has been recorded during the Tertiary. Additionally, the cooling/ exhumation path for the structurally lowermost Pelagonian parts is dated during the Oligocene-Miocene [38,39,47,48,86,155,156].
About the geotectonic position of the Pelagonian basement, it is regarded: I. either as the Mesozoic eastern passive margin of the Apulia continent, with a main wide ocean basin in the east, so-called the Neothethyan Meliata/Maliac-Axios/Vardar ocean basin [6,11,13,43,154,157,158] or II. as a microcontinent emerging in the middle of two separate ocean basins that were operating more or less contemporaneously during the Alpine orogeny; these were the Pindos ocean to the west and the Axios/Vardar ocean to the east east [3,113,115,116,159,160,161,162]. As accruing in these interpretations, the ophiolite rocks on both the western and eastern parts of the Pelagonian nappe should have originated either from one or two ocean sources, respectively. This leads to an ongoing discussion about the direction of the ophiolite`s obduction onto the Pelagonian continent during the Middle-Late Jurassic; either only one main W- to SW-ward obduction direction (e.g., [1,6,11,13,20,37,136,158,163,164]) or both W- to SW-ward and E- to NE-ward (e.g., [3,113]) or even only E- to NE-ward (e.g., [159,162]). Additionally, the last view sees only one main ocean basin, the Pindos basin, West of the Pelagonia.
Figure 10. Geological-structural map and representative cross-section of the eastern tectonic boundary of the Axios/Vardar zone with the Serbo-Macedonian/Rhodope massif. The Pelagonian nappe is sandwiched between Europe (the Serbo-Macedonian/Rhodope nappes pile) and Apulia (the External Hellenides). (Based on [119,120,149,150]). Abbreviations as in Figure 3.
Figure 10. Geological-structural map and representative cross-section of the eastern tectonic boundary of the Axios/Vardar zone with the Serbo-Macedonian/Rhodope massif. The Pelagonian nappe is sandwiched between Europe (the Serbo-Macedonian/Rhodope nappes pile) and Apulia (the External Hellenides). (Based on [119,120,149,150]). Abbreviations as in Figure 3.
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The Sub-Pelagonian Zone is composed of a sequence of Triassic to Jurassic pelagic carbonate and siliceous sediments, which lie tectonically over the Mesozoic platform carbonate sediments of the Pelagonian nappe. Today, it is traced along the western Pelagonian margin. The characteristic phase of the Sub-Pelagonian zone is the red, ammonite-bearing, pelagic sediments of the Hallstatt phase of the Late Triassic age (Figure 12). Ophiolites and ophiolitic mélanges overthrust the Sub-Pelagonian deep-water sediments (Figures 3, 4). The latter emplacement took place during the Mid-Late Jurassic, following the general geotectonic history of the Alpine orogeny in the Hellenides.
However, the Sub-Pelagonian zone remains an under-debate zone concerning its paleogeographic existence and geotectonic setting. One scenario wants the Sub-Pelagonian zone to be the Mesozoic continuation of the western Pelagonian continental margin to the continental slope and the deeper basin area towards the Pindos ocean basin (e.g., [3,19,113,165]). The opposing theory of only one wide ocean basin to the eastern of the Pelagonian nappe (i.e., Meliata/Maliac-Axios/Vardar ocean basin) explains the position of the Sub-Pelagonian zone along the western Pelagonian side as such of a tectonic nappe nature, having been thrust, together with the ophiolite belt, from the east to the west on the Pelagonian nappe [6,13,20,24,43,136,154]. In any case, the Sub-Pelagonian zone was affected by the Late Jurassic deformation and the subsequent, younger deformational events recognized in the Hellenides orogenic belt, placing the Sub-Pelagonian zone within the Internal Hellenides zones.
Figure 11. a. and b. P/T/t Alpine tectono-metamorphic path and exhumation history: (a) of the Pelagonian and (b) Paikon basements; Red color: granite intrusions in the Pelagonian basement [6,133,142]). c. Meso- and microscale features of the deformational events: I. Main S1 foliation related to isoclinal recumbent folds. Asymmetric F2 folds overprint the previous isoclinal folds. S2 foliation is also shown. Garnet mica schist (Upper Pelagonian segment, Peternik unit; II. Recrystallization of sericite along S2 planes around white mica porphyroclast. The asymmetry of the mica fish indicates a top-to-NW sense of shear during D2. XZ section. Garnet-bearing mica gneiss of the Upper Pelagonian segment, Peternik unit; III. Garnet with characteristic zonation and internal fabric (Si) rotated during D2. Chlorite aggregates (after garnet) and sericite growth in asymmetric pressure shadows and along the dominating S2 foliation planes. Garnet σ-clasts and S-C fabric indicate a top-to-NW sense of shear. XZ section. Garnet mica schist (Lower Pelagonian segment); IV. S-C fabric in mica gneiss from the tectonic contact between the lower and upper duplicated Pelagonian parts during the Late Jurassic. X-Z section. Sense of shear top-to-WNW; V, VI. Augengneisses of the Pelagonian tectonic sheets in the Axios/Vardar zone (Livadia and Peternik units). Feldspar σ-clasts, S-C fabrics, and shear bands indicate a top-to-NE sense of shear (D3 event). XZ section; VII. S1/S2 relationship. S1 is defined by white mica and chloritoid. Chloritoid rotates into S2. YZ section. Garnet-chloritoid mica schist (Lower Pelagonian segment); VIII. Shear bands and S-C fabric indicate a top-to-WNW sense of shear during D2. Intensive chloritization of D1 garnet σ-clasts took place during D2. X-Z section. Garnet mica schist (Lower Pelagonian segment); IX. D1 and D2 fold realms on the Triassic-Jurassic carbonate sequence of the Paikon subzone (Gandatch marbles). Isoclinal, recumbent folds related to the D1 event are overprinted by the D2 event. The S2 foliation dominates; X. Calcite σ-clast in the Triassic-Jurassic carbonate sequence of the Paikon basement (Gandatch marbles), indicating a top-to-SW sense of movement (D1); XI. S1/S2 fabric in the Late Jurassic volcanosedimentary formation of the Paikon subzone (Kastaneri formation). The old S1-foliation is strongly reoriented along the S2 foliation, forming a granulation cleavage fabric. Due to the strong translation along the S2-planes, the two foliations are usually developed parallel to one another, so that only one fabric element seems to be recognized on the geological formation and this is the S2-foliation. Paikon subzone, Axios/Vardar zone; XII. Brittle D4 thrust zone towards SW, cutting the main S2 foliation. Brittle also is the D5 semi-low angle normal fault zone cutting the D4 thrust zone and S2 foliation with a top-to-NE sense of shear. Late Jurassic volcanosedimentary formation of the Paikon subzone (Kastaneri formation); XIII, XIV. B-axis scattering of D2-isoclinal folds due to their re-orientation subparallel to the X-axis of the strain ellipsoid. D2 stretching lineation SW-ward plunging. Triassic-Jurassic carbonate sequence of the Paikon basement (Gandatch marbles); XV. Kink-folds in the Late Jurassic-Early Cretaceous carbonate Griva formation of the Paikon subzone related to the Paleocene-Eocene compressional event (D4). Microscopic pictures: III, IV, VII and VIII with one Nicol, II, VI with crossed Nicols (modified after [6,142]).
Figure 11. a. and b. P/T/t Alpine tectono-metamorphic path and exhumation history: (a) of the Pelagonian and (b) Paikon basements; Red color: granite intrusions in the Pelagonian basement [6,133,142]). c. Meso- and microscale features of the deformational events: I. Main S1 foliation related to isoclinal recumbent folds. Asymmetric F2 folds overprint the previous isoclinal folds. S2 foliation is also shown. Garnet mica schist (Upper Pelagonian segment, Peternik unit; II. Recrystallization of sericite along S2 planes around white mica porphyroclast. The asymmetry of the mica fish indicates a top-to-NW sense of shear during D2. XZ section. Garnet-bearing mica gneiss of the Upper Pelagonian segment, Peternik unit; III. Garnet with characteristic zonation and internal fabric (Si) rotated during D2. Chlorite aggregates (after garnet) and sericite growth in asymmetric pressure shadows and along the dominating S2 foliation planes. Garnet σ-clasts and S-C fabric indicate a top-to-NW sense of shear. XZ section. Garnet mica schist (Lower Pelagonian segment); IV. S-C fabric in mica gneiss from the tectonic contact between the lower and upper duplicated Pelagonian parts during the Late Jurassic. X-Z section. Sense of shear top-to-WNW; V, VI. Augengneisses of the Pelagonian tectonic sheets in the Axios/Vardar zone (Livadia and Peternik units). Feldspar σ-clasts, S-C fabrics, and shear bands indicate a top-to-NE sense of shear (D3 event). XZ section; VII. S1/S2 relationship. S1 is defined by white mica and chloritoid. Chloritoid rotates into S2. YZ section. Garnet-chloritoid mica schist (Lower Pelagonian segment); VIII. Shear bands and S-C fabric indicate a top-to-WNW sense of shear during D2. Intensive chloritization of D1 garnet σ-clasts took place during D2. X-Z section. Garnet mica schist (Lower Pelagonian segment); IX. D1 and D2 fold realms on the Triassic-Jurassic carbonate sequence of the Paikon subzone (Gandatch marbles). Isoclinal, recumbent folds related to the D1 event are overprinted by the D2 event. The S2 foliation dominates; X. Calcite σ-clast in the Triassic-Jurassic carbonate sequence of the Paikon basement (Gandatch marbles), indicating a top-to-SW sense of movement (D1); XI. S1/S2 fabric in the Late Jurassic volcanosedimentary formation of the Paikon subzone (Kastaneri formation). The old S1-foliation is strongly reoriented along the S2 foliation, forming a granulation cleavage fabric. Due to the strong translation along the S2-planes, the two foliations are usually developed parallel to one another, so that only one fabric element seems to be recognized on the geological formation and this is the S2-foliation. Paikon subzone, Axios/Vardar zone; XII. Brittle D4 thrust zone towards SW, cutting the main S2 foliation. Brittle also is the D5 semi-low angle normal fault zone cutting the D4 thrust zone and S2 foliation with a top-to-NE sense of shear. Late Jurassic volcanosedimentary formation of the Paikon subzone (Kastaneri formation); XIII, XIV. B-axis scattering of D2-isoclinal folds due to their re-orientation subparallel to the X-axis of the strain ellipsoid. D2 stretching lineation SW-ward plunging. Triassic-Jurassic carbonate sequence of the Paikon basement (Gandatch marbles); XV. Kink-folds in the Late Jurassic-Early Cretaceous carbonate Griva formation of the Paikon subzone related to the Paleocene-Eocene compressional event (D4). Microscopic pictures: III, IV, VII and VIII with one Nicol, II, VI with crossed Nicols (modified after [6,142]).
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In addition, the Triassic-Eocene Parnassos carbonate unit develops in central Greece (Figures 1, 2), terminating to an Eocene-Oligocene flysch and known for the rιch bauxite deposits. It is regarded as a platform carbonate sequence that was possibly deposited on the more western Pelagonian parts. On the contrary, due to the absence of a clear stratigraphic gap in its tectonostratigraphic column, from the Triassic to the Eocene-Oligocene, the Parnassos unit is considered to be a part of the External Hellenides zones, forming a reef coralgal build-up belt near the Pelagonian western margin [125,166,167,168,169]. In any case, the geotectonic and paleogeographic setting of the Parnassos carbonate sequence remains under debate today.
The Parnassos unit, together with the Pelagonian nappes, overthrust W-ward the External Hellenides Pindos zone during the Paleocene-Eocene. Its continuation is recognized further to the North in the ``high karst`` zone of the Croatian and Bosnia-Herzegovina in the Dinarides, where important bauxite deposits are also appearing, as they are in the Parnassos unit in Greece (Figures 1, 2; [125,167,168,169,170]).

2.2.2. Axios/Vardar Zone

The Axios/Vardar zone is structurally another very complicated Hellenides’ zone composed of units of both, continental and oceanic origin. The whole zone forms a typical Tertiary thrust and folds belt, where ophiolites and Mid-Late Jurassic ophiolitic mélanges are intensively imbricated with Paleozoic continental basement rocks and low-grade to non-metamorphic Mesozoic sedimentary series, as well as Late Jurassic arc-type calc-alkaline granitoid cross-cutting in places either basement slivers or ophiolites (e.g., Fanos granite, Figures 4, 13, 14; [30,164,171,172]). The Axios/Vardar zone is traditionally subdivided into three (3) subzones. From the West to the East, they are the Almopia, the Paikon and the Paionia subzones (Figures 3, 4, 13, 14; [27,37]).
Several different interpretations have been proposed about the geotectonic position and structural evolution of the Axios/Vardar zone. Some authors believe that there were two ocean basins, separated by the continental Paikon volcanic arc. Western of the Paikon arc there was the Almopia ocean basin with Triassic oceanic crust, while eastern of the
Figure 12. Field photographs of the ?Sub-Pelagonian zone at the western margin of the Pelagonian nappe (modified after Kilias et al. 2016): I. Green schists and deep-water sediments of the Sub-Pelagonian zone with a conjugate set of tension gashes related to a vertical maximum σ1-axis and a subhorizontal σ3-axis, a dynamics that coincides with the Oligocene-Miocene extension (D5 event) and the carbonate Olympos-Ossa unit exhumation under the downwardly detached blue schists unit and the Pelagonian nappe pile; II, III, IV, V. Multicoloured deep-water sediments composed of green schists, red- and green-coloured radiolarian cherts, pelagic red carbonates and metapelite rocks, possibly of Jurassic age. They show low-grade metamorphism, but they are intensively deformed by compressional structures (e.g., isoclinal folds, reverse shear bands, and brittle thrust faults), as well as extensional structures (e.g., down-dip shear bands, S-C fabric and semi-ductile normal fault zones). These deep-sea sediments, together with the ophiolite belt, overthrust the neritic Triassic-Jurassic carbonate rocks of the Pelagonian nappe during the Middle-Late Jurassic. The sense of movement during their initial emplacement is not clear here due to the intensive after-emplacement multi-phase deformation that affected these rocks; VI. Red carbonate sediments lying under the previous multicoloured metasediments. They represent a deep-sea basin environment and show sedimentary brecciation as well as turbidity layering. Their contact with the underlying Paleozoic Pelagonian schists and Triassic-Jurassic Pelagonian platform carbonate cover is a Neogene-Quaternary normal fault zone. The age of this sequence remains under debate. It should most likely be of Jurassic age equivalent to the previously described multicoloured deep-water sediments of the ?Sub-Pelagonian zone, having also overthrust the neritic Triassic-Jurassic Pelagonian platform carbonate cover during the Middle-Late Jurassic together with the Neotethyan ophiolites.
Figure 12. Field photographs of the ?Sub-Pelagonian zone at the western margin of the Pelagonian nappe (modified after Kilias et al. 2016): I. Green schists and deep-water sediments of the Sub-Pelagonian zone with a conjugate set of tension gashes related to a vertical maximum σ1-axis and a subhorizontal σ3-axis, a dynamics that coincides with the Oligocene-Miocene extension (D5 event) and the carbonate Olympos-Ossa unit exhumation under the downwardly detached blue schists unit and the Pelagonian nappe pile; II, III, IV, V. Multicoloured deep-water sediments composed of green schists, red- and green-coloured radiolarian cherts, pelagic red carbonates and metapelite rocks, possibly of Jurassic age. They show low-grade metamorphism, but they are intensively deformed by compressional structures (e.g., isoclinal folds, reverse shear bands, and brittle thrust faults), as well as extensional structures (e.g., down-dip shear bands, S-C fabric and semi-ductile normal fault zones). These deep-sea sediments, together with the ophiolite belt, overthrust the neritic Triassic-Jurassic carbonate rocks of the Pelagonian nappe during the Middle-Late Jurassic. The sense of movement during their initial emplacement is not clear here due to the intensive after-emplacement multi-phase deformation that affected these rocks; VI. Red carbonate sediments lying under the previous multicoloured metasediments. They represent a deep-sea basin environment and show sedimentary brecciation as well as turbidity layering. Their contact with the underlying Paleozoic Pelagonian schists and Triassic-Jurassic Pelagonian platform carbonate cover is a Neogene-Quaternary normal fault zone. The age of this sequence remains under debate. It should most likely be of Jurassic age equivalent to the previously described multicoloured deep-water sediments of the ?Sub-Pelagonian zone, having also overthrust the neritic Triassic-Jurassic Pelagonian platform carbonate cover during the Middle-Late Jurassic together with the Neotethyan ophiolites.
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Paikon arc evolved in a back-arc region the Paionia ocean basin during the Mid-late Middle Jurassic, simultaneously with the subduction of the Almopia ocean beneath the Paikon continental arc [16,22,23,113,160,161,173]. The Middle-late Middle Jurassic age of the Paionia oceanic crust is also documented by [174] using radiolarian ages of deep-sea sediments covering the Paionia oceanic crust. Furthermore, the same Middle-late Middle Jurassic age for the Paionia oceanic crust is also concluded by [175] who determined an isotopic age of 166 Ma for a plagiogranite from the Paionia ophiolites. Similar Middle Jurassic ages (Bajosian) of radiolarian assemblages in the cherts stratigraphically on top of the Vourinos ophiolites at the western edge of the Pelagonian nappe (Figures 3, 14, 15) have also been recorded by [176].
In another view, the Paionia ocean basin here is equivalent to the Axios/Vardar ocean basin, formed during the Mid-late Middle Jurassic in a supra-subduction zone setting or back-arc region, behind an ensimatic island arc, that resulted from an Early-Middle Jurassic intra-oceanic subduction in the Neotethyan Meliata/Maliac ocean basin [11,20,135,175,177]. In this case, the Paikon subzone was interpreted as a tectonic window of Pelagonian origin, where both the ophiolites of the Almopia’s and Paionia’s subzones, together with the ensimatic arc volcano-sedimentary products, were W-SW-ward obducted during the Mid-Late Jurassic (Callovian-Oxfordian) from a single ocean basin and it was the Neotethyan Axios/Vardar basin located East of the Pelagonian continental margin (Figures 3, 4, 13, 14; [2,6,13,135,142,164,178]). On the other hand, ophio- lites of the eastern-northeastern part of the single Axios/Vardar ocean basin were obducted simultaneously during the Mid-Late Jurassic on the European margin but with an N-NE-ward sense of movement (Figures 4, 10, 13, 14; [24,41,179,180,181,182]). In this case, the simultaneous during the Mid-Late Jurassic SW-ward and NE-ward ophiolites obduction on both continental margins of the Pelagonia and Europe, respectively, was related to an arcuate type NW-ward convex intra-oceanic subduction zone of the western-northwestern more part of the Neotethyan, the Meliata/Maliac ocean realm [17,24].
Figure 13. Geological-structural map and representative cross-section of the whole Axios/Vardar zone (including the Almopia, Paikon and Paionia subzones) and its contact with the eastern Pelagonian margin at Voras and Paikon/Tzena Mts. The Paikon basement and its Triassic-Jurassic carbonate cover are shown as a tectonic window of Pelagonian origin beneath the obducted Neotethyan Axios/Vardar ophiolites and the Mid-Late Jurassic island arc-type magmatic products (Volcanosedimentary series, VS). The Late Jurassic-Early Cretaceous and Late Cretaceous sedimentary sequences are also shown (based on [6,27,142,143,160,161]). Abbreviations as in Figure 3.
Figure 13. Geological-structural map and representative cross-section of the whole Axios/Vardar zone (including the Almopia, Paikon and Paionia subzones) and its contact with the eastern Pelagonian margin at Voras and Paikon/Tzena Mts. The Paikon basement and its Triassic-Jurassic carbonate cover are shown as a tectonic window of Pelagonian origin beneath the obducted Neotethyan Axios/Vardar ophiolites and the Mid-Late Jurassic island arc-type magmatic products (Volcanosedimentary series, VS). The Late Jurassic-Early Cretaceous and Late Cretaceous sedimentary sequences are also shown (based on [6,27,142,143,160,161]). Abbreviations as in Figure 3.
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In a more recent scenario, the “maximum allocthony hypothesis” suggests that the Axios/Vardar zone is allochthonous, emplaced secondary as a tectonic nappe in its present outcropping position, and the Axios/Vardar suture zone is located at depth along the NE boundary of the Rhodopes, building up the root zone of the most eastern nappes pile of the Internal Hellenides (e.g., [17,24,119,120,183]). Continent collision of Europe and Pelagonia as a fragment of Apulia, intense imbrication and nappes stack, following up the finale closure and suturing of the Middle-Late Jurassic Axios/Vardar ocean basin, occurred in the Paleogene (Paleocene-Eocene) after subduction during the Late Cretaceous of the remnants of the Axios/Vardar ocean realm beneath the European continental margin [17,19,21,24,59,113,117]. This Axios/Vardar suturing is continued further to the North, called the Sava zone and to the East, to the Ankara suture zone in Turkey (Figures 1, 2; [11,21,24,113,184,185,186]).
Furthermore, slices of Paleozoic basement rocks, gneissics and schists (e.g., Peternik and Livadia units), tectonically emplaced in between the Axios/Vardar Mesozoic sequences, form the root parts of the Upper Pelagonian unit (KoWPZ) that overthrust the Lower Pelagonian unit (EPZ). It was caused by internal Middle-Late Jurassic thrusting that cut the Pelagonia into two main segments, of which one was placed upon the other, with the Upper being rooted in the Axios/Vardar zone (Figures 2, 3, 13; [6,187]).

2.2.3. Cycladic massif

The Cycladic massif is composed of a complicated, heterogeneous nappe stack system, including parts of the Internal and External Hellenides zones. In a general view, the Cycladic tectonostratigraphy can be given as follows, although it cannot be mapped entirely in a region due to the evolution of individual parts of the whole Cycladic massif section in the small islands of the central Aegean sea (Cyclades islands), the Attica peninsula and the south Evia island (Figures 1, 2: [45,80,88,89,90,91,92,93,94,95,98,110,188,189]). The deepest unit comprises a Variscan or older basement of schists and gneisses, as well as Carboniferous granitoid (~300 Ma) and abundant Oligocene-Miocene migmatites and granites. It constitutes the most deeply exhumed parts of the Hellenides belonging to the Apulian basement of the External Hellenides, possibly equivalent to the Menderes Massif, as discussed previously [9,98,188]. The Cycladic basement is overlain by a post-Variscan, Alpine sedimentary cover, containing marbles, metapelites and Early Triassic volcanic intercalations, which form the metamorphic continuation of the Apulian passive margin, equivalent to the Gavrovo carbonate platform sequence on the Hellenic mainland. The marbles contain emery and metabauxites in some places, e.g., in the Naxos island (Figures 4, 7, 15; [9,188,190]). Tectonically, on the Alpine passive margin sequence rests the Paleocene-Eocene high-pressure belt of marbles, metapelites and metabasites intercalations, as in the Olympos-Ossa and Pelion areas, but here in the Cyclades, with eclogites also occurrences. Highly attenuated high-pressure also metamorphosed ophiolitic mélanges containing slivers of oceanic crust formed during ~80-70Ma [191,192] are usually incorporated in this Paleocene-Eocene high-pressure belt. They are considered the NE-ward subducted parts of a small Pindos/Cyclades oceanic crust beneath the Pelagonian nappe [17,24,193]. Furthermore, all lithologies from the basement to the post-Variscan sedimentary cover show this Paleocene-Eocene high-pressure metamorphic overprint [45,46,49,91,189]). Additionally, an intensive Oligocene-Miocene high-temperature/low-pressure (HT/LP) retrogressive metamorphism until migmatization and granitization is recognized elsewhere in the Cyclades massif related to extension and crustal uplift [88,95]. The P/T/t tectonometamorphic path of the Cycladic massif is given in detail in Figure 8b.
On the Cycladic high-pressure belt, emplaced tectonically during the Paleocene-Eocene, the Pelagonian nappe with the Mid-Late Jurassic obducted Neotethyan ophiolites. Today, due to the intensive Aegean Oligocene-Miocene extension, they were tectonically denudated and eroded and only small rests are saved or as conglomerate clasts in the sediments of the Neogene basins [80,138,139].
The Cycladic high-pressure belt constitutes a typical metamorphic core complex exhumed during the Oligocene-Miocene extension in Hellenides along normal detachment faults associated with mylonites formation and high-temperature metamorphism in the deeper tectonic levels [9,10,89,90,91,92,94,95,110,111,188,194]. Nevertheless, several studies argue that extension started no earlier than the Early Miocene [90,94,110,188,194].
Figure 14. Geological-structural map and representative cross-section of the eastern Axios/Vardar zone (Paionia subzone) and the Circum-Rhodope belt, which is regarded as the sedimentary sequence initially deposited on the western margin of the Serbo-Macedonian massif (European margin). The Fanos granite, connected with an ensimatic island-arc, is shown overthrust together with the ophiolites on the eastern Pelagonian margin. The tectonic contact between Europe (Serbo-Macedonian) and Apulia (Pelagonian) is also shown (based on [119,120,164,183,256,257]). Abbreviations as in Figure 3.
Figure 14. Geological-structural map and representative cross-section of the eastern Axios/Vardar zone (Paionia subzone) and the Circum-Rhodope belt, which is regarded as the sedimentary sequence initially deposited on the western margin of the Serbo-Macedonian massif (European margin). The Fanos granite, connected with an ensimatic island-arc, is shown overthrust together with the ophiolites on the eastern Pelagonian margin. The tectonic contact between Europe (Serbo-Macedonian) and Apulia (Pelagonian) is also shown (based on [119,120,164,183,256,257]). Abbreviations as in Figure 3.
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2.2.4. Serbo-Macedonian and Rhodope metamorphic massifs

The Serbo-Macedonian and Rhodope metamorphic massifs are located in the northeastern Hellenides region and they continue with the same composition and structural architecture farther North, in Bulgaria and Serbia and perhaps farther East in northwestern Turkey (Figures 1, 2, 3, 4; [9,12,18]). They are composed of a complex Alpine nappes stack of several metamorphic units or terranes of both continental and oceanic origin, with the Rhodopes constituting the core of the arc-type Hellenic orogenic belt (Figures 1, 2). The Axios/Vardar zone and the Circum-Rhodope belt border the Serbo-Macedonian/Rhodope metamorphic province to the West and the Maritza dextral strike-slip fault in Bulgaria forms its other boundary to the Northeast (Figures 1, 2, 3, 4).
Figure 15. Geological-structural map and representative cross-section of the Vourinos ophiolites and the western Pelagonian margin, including the Zygosti stream ophiolite rocks and the Late Jurassic-Early Cretaceous and Late Cretaceous carbonate series, unconformably overlying either the Vourinos ophiolites or the Zygosti stream ophiolite rocks (based on [25,116,145]). Abbreviations as in Figure 3.
Figure 15. Geological-structural map and representative cross-section of the Vourinos ophiolites and the western Pelagonian margin, including the Zygosti stream ophiolite rocks and the Late Jurassic-Early Cretaceous and Late Cretaceous carbonate series, unconformably overlying either the Vourinos ophiolites or the Zygosti stream ophiolite rocks (based on [25,116,145]). Abbreviations as in Figure 3.
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The various Serbo-Macedonian and Rhodope nappes were successively emplaced one on top of the other by intense compressional tectonics and thrusting from the Jurassic to the Tertiary. In any case, compressional tectonics and thrusting are related to multiple subduction processes, high- to ultra-high-pressure metamorphism and the final closure of the Meliata/Maliac-Axios/Vardar ocean basin, which developed, as was already mentioned, in the peculiar position between the Apulia plate to the West and the European plate to the East. Today, the tectonic contacts between the several nappes remain only partly as the original thrust faults related to the orogeny nappes stack. They have usually been reworked to Tertiary low angle, normal detachment faults due to extensional tectonics, following progressively, the several compressional stages and nappes stacking. This Tertiary extensional period was also accompanied by high-grade metamorphism, syn- to post-tectonic granitoid intrusions and abundant migmatite formations (Figures 3, 4; e.g., [4,7,17,18,24,40,43,114,154,195,196,197,198,199]).
The Serbo-Macedonian and Rhodope metamorphic nappe piles in the Greek part are made up of the tectonically lower to the tectonically higher nappes as follows: in the Rhodope province by the Pangaion, Sidironero and Kimi units and in the Serbo-Macedonian province by the Kerdylia and Vertiskos units (Figures 3, 4, 5).

2.2.4.1. Rhodope massif

  • I. Pangaion unit
The structurally lowermost Rhodope Pangaion unit consists of mica-schists, gneisses and marble intercalations, covered in the higher levels by a thick marble sequence (Figures. 5, 16). The age of the unit is under debate. In recent works, it das been described as a segment of Variscan or older continental crust covered by a thick marble sequence, of possible Mesozoic age [17,22,23,24,195,200,201]. Nevertheless, other authors suggest a Paleozoic age for the whole Pangaion unit, including the overlain thick marble sequence [202,203,204,205,206]. The Pangaion unit was intruded by Oligocene-Miocene granitoids and during the Tertiary (Oligocene-Miocene) it underwent low-grade metamorphism under greenschist facies conditions and an extensional regime [114,195,207]. Nevertheless, some authors also describe a possible Paleocene-Eocene high-pressure metamorphic event for the lower Rhodope Pangaion unit (45,8 ± 5,8Ma; [57,208]).
The Pangaion unit is bordered to the Northeast by the Nestos shear zone (Figures 3, 4, 16). It is interpreted as a ductile thrust zone along which the tectonically middle Rhodope Sidironero unit overthrusted the Pangaion unit during the Tertiary (Paleocene-Eocene; Jawn-Awe et al. 2010, Froitzheim et al. 2014), although some other authors date the Nestos thrust zone as of Cretaceous or Jurassic age (e.g., [18,22,23,196]). Furthermore, [201] interpreted the Nestos thrust zone as a suture zone between the lower Rhodope Pagaion unit and the middle Rhodope Sidironero unit of Jurassic-Cretaceous age.
On the contrary, the southwestern border of the Pangaion unit is controlled by an Oligocene-Miocene normal detachment zone along which the Serbo-Macedonian massif was detached SW-wards, resulting in the final exhumation of the Pangaion metamorphic core complex and the formation of the supra-detachment Neogene Strymon basin (Figures 3, 4, 16; [7,114,195,209,210]).
The Pangaion unit is variably interpreted until today as an Oligocene-Miocene metamorphic core complex belonging to the External Hellenides Apulia plate [17,24,114,119,120,183] or a microcontinent, named Drama or Thracia terrane [4,201] accreted to the European continent, either during the Early-Middle Jurassic following the closure of a Triassic age`s ocean basin [22,23] or during the Cretaceous [18,196,201]. Furthermore, [211] interpret the Pangaion unit as a continental crust derived from the Pelagonian segment.
  • II. Sidironero unit
The Sidironero unit tectonically overlies the Pangaion unit and is composed of rocks of oceanic and continental origin. We distinguish amphibolites, ortho- and paragneisses, as well as marbles, migmatites and metaophiolites. The orthogneises are derived from Late Jurasic-Early Cretaceous arc-type granitoids as well as from Variscan granitoids that intruded the older Paleozoic Sidironero basement rocks [201,212,213]. The Late Jurasic-Early Cretaceous arc type granitoid intrusions highlight the Sidironero unit as a Late Jurassic subduction-related volcanic arc and the Pangaion unit as a microcontinent underthrust possibly the Sidironero unit during the Late Jurassic, although this interpretation remains under discussion (e.g., [17,22,23,24,201,205,206]). Furthermore, Eocene-Oligocene adakitic and Oligocene-Miocene calc-alkaline granitoids intrude also the basement rocks during an extensional tectonic regime (Figures 3, 4, 5, 16; [24,114,197,214,215,216,217,218,219,220]). Metaophiolites occuring in between the Sidironero continental rocks yielded a ca. 160 Ma age for their protolith. They have been interpreted as equivalent to the Axios/Vardar ophiolites belt, emplaced tectonically in the Sidironero unit during the Late Cretaceous-Paleocene following the closure of the Axios/Vardar oceanic basin [17,24].
In the Greek and southern Bulgaria Rhodope province, two dome-shape tectonic windows, the Kesebir-Kardamos and Kechros domes, exhumed along Tertiary normal detachment faults below the upper Rhodope Kimi unit and composed by Paleozoic gneissic and schists rocks intruded by Variscan and Triassic, as well as Tertiary granitoids, have been grouped with the lowermost Pangaion metamorphic core complex (Figures 3, 4; [17,18,40,201]). Nevertheless, taking into account that their composition and structural evolution are not exactly defined, some questions about their geotectonic setting as equivalent to the Pangaion metamorphic core complex are raised. So, the Kesebir-Kardamos and Kechros domes, are not characterized by the thick carbonate cover sequence dominated in the Pangaion unit and they were affected by a higher- grade Tertiary metamorphism than the Pangaion unit associated with migmatization that is not observed in the Pangaion unit [40,195,221,222,223]. Furthermore, they show cooling/exhumation path during Eocene-Oligocene extension (~42-35 Ma), which is the same as the Sidironero unit (Figures 4, 17a; [225]). The cooling/exhumation path for the Pangaion unit was related to Oligocene-Miocene extension (26-10 Ma, Figures 4, 17a; [224,225]). According to that, if we also consider the structural position of the Kesebir-Kardamos and Kechros domes, exactly below the Kimi unit, their geotectonic-paleogeographic regime as equivalent to the Pangaion unit is under dispute.
Figure 16. Geological-structural map and representative cross-section of the Pangaion metamorphic core complex and the Serbo-Macedonian/Rhodope nappes pile. The Paleocene-Eocene Nestos thrust zone [17,24] and the Oligocene-Miocene Strymon valley normal detachment fault zone [114,210], as well as the Miocene-Pliocene supradetachment Strymon basin, are shown [7,30]. Abbreviations as in Figure 3.
Figure 16. Geological-structural map and representative cross-section of the Pangaion metamorphic core complex and the Serbo-Macedonian/Rhodope nappes pile. The Paleocene-Eocene Nestos thrust zone [17,24] and the Oligocene-Miocene Strymon valley normal detachment fault zone [114,210], as well as the Miocene-Pliocene supradetachment Strymon basin, are shown [7,30]. Abbreviations as in Figure 3.
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High-pressure eclogite facies metamorphic paragenesis remnants and ultrahigh-pressure diamond-bearing rocks, including mainly basic Morb-type rock bodies, were described for the Sidironero unit. Isotopic analysis data for the high-pressure rocks have given various ages that have been partlially interpreted differently by the several authors. In any case, high-pressure metamorphic conditions were related to subduction, compression and crustal thickening due to the thrusting or crustal sinking due to overloading setting of the nappes stack. The ages of the recognized high-to ultrahigh-pressure metamorphic events are: ca. 200 Ma [54,55,56,117,201]; 150-140 Ma [53,223]; 92,7-70,5 Ma [58,59]; and 51-45 Ma [51,53,57,226]. The high-pressure metamorphic events were finally strongly reworked by an Eocene-Oligocene retrogressive high-temperature in the amphibolite facies metamorphism, partly reaching migmatization conditions and granitoιd genesis. This high-temperature metamorphic event was related to extension and crustal exhumation. Moreover, it is the dominant metamorphism in the Sidironero unit and strongly reworked all other older metamorphic events [51,53,114,195,197,207]. The pre-Paleocene-Eocene ages of the high- and ultrahigh-pressure metamorphic events described for the Sidironero metamorphic rocks have also been detected in the tectonically overlying upper Rhodope Kimi. These high-pressure rocks were interpreted as parts of the Kimi unit that were imbricated in between the Sidironero rocks due to younger, successively thrusting events [24,59].
Furthermore, a migmatization event of ca. 140 Ma for the Sidironero unit referred to by [223], remains without any clear explanation about its geotectonic framework. In any case, the two migmatization ages at the Early Cretaceous and Eocene indicate the multiphase evolution of the Alpine orogeny in the Rhodope massif.
  • III. Kimi unit
The Kimi unit represents the upper tectonic nappe of the Rhodope nappes stack. It is composed of various metamorphic rocks continental origin (ortho- and paragneisses, amphibolites, thin marble layers, gneiss-schists) and meta-ophiolites. The protolith ages of the orthogneisses were dated as Paleozoic (Ordovician and Silurian), while the meta-ophiolites yielded Early Paleozoic and Triassic protolith ages [41,213,227,228,229,230].
High- to ultrahigh-pressure metamorphic rocks, including eclogites and diamond-bearing gneisses, were also recognized in the upper Rhodope Kimi unit. Geochronological isotopic data yielded again here a large dispersion of ages for the high- to ultrahigh metamorphic rocks and different interpretations for their development. The isotopic ages are described as follows: ca. 200 Ma, ca. 158 Ma, ca. 125 Ma and 82-74 Ma [50,59,117,231,232,233]. The 200 Ma age [117,231] was related to the diamond occurrence in gneisses and meta-ophiolites. This event has been possibly related to the closure of the Paleotethys around 200 Ma ago (in the Early Jurassic) and thus represents the palaeogeographic location of the actual suture of the Paleotethyan ocean [24,56,117,231]. The high-pressure metamorphism that occurred during the Late Cretaceous (~80-85 Ma; [59,117,232,233]) is in a very good concordance with the proposed Late Cretaceous subduction of the remnants of the Neotethyan Axios/Vardar Ocean below the European margin and the formation of a Late Cretaceous magmatic arc with arc-type granitoid inclusions in the European margin (Sredna-Gora massif; [59,213,234,235,236]). For the other high-pressure events referred to, there are still no clear evidences for their structural evolution or geotectonic regime.
The Late Cretaceous high-pressure metamorphic event, associated with compression and nappe stacking, was progressively reworked by a Paleocene-Eocene retrogressive high-temperature in the amphibolite facies metamorphism, partly reaching migmatization and granitization conditions. High-grade temperature metamorphism was related to extension, tectonic denudation of the Rhodope nappe stack and crustal exhumation [7,17,24,40,67,198,230].
Moreover, regarding the cooling/exhumation paths of the Kimi, Sidironero and Pangaion Rhodope units, the whole Rodope nappes stack is characterized by a gradual cooling/exhumation history during extension. From the top to the bottom units it is: The cooling/exhumation path for the upper Kimi unit was dated at the Paleocene-Eocene (~60-48 Ma), for the middle Sidironero unit at the Eocene-Oligocene (~42-30 Ma) and finally for the lowermost Pangaion unit at the Oligocene-Miocene (~26-10 Ma). It shows the progressive exhumation evolution of the Rhodope units from the top to the bottom. (Figures 4, 17a; [7,17,24,40,67,114,195,197,199,207,225]).
Finally, we agree with the interpretation that the upper Rhodope Kimi nappe represents the Mesozoic western continental margin of Europe, where the Neotethyan ophiolites were obducted NE-ward during the Mid-Late Jurassic arc-continent collision [24,41,119,120].

2.2.4.2. Serbo-Macedonian massif

  • I. Kerdylia unit
As already mentioned, the Serbo-Macedonian massif is divided into the lower Kerdylia and the higher Vertiskos units that form parts of a nappe stack (Figures 3, 4, 5). The Kerdylia unit is composed of Paleozoic ortho- and paragneisses and schists, as well as granitoid intrusions and a relatively thin marble cover of unknown age, constituting the highest lithostratigraphic sequence of the unit (Figure 16). [22,23] records the marble cover as a possible Triassic sedimentary series, deposited on a passive continental margin. Migmatites also occur with their age until today under discussion, possibly ranging from the Paleozoic to the Tertiary [1,68,237,238]. The Kerdylia unit was tectonically placed secondary on top of the western side of the Rhodope Pangaion unit along an Oligocene-Miocene normal detachment fault zone related to the final exhumation of the lowermost Rhodope Pangaion unit (Figures 3, 4, 16; [114,118,195,207,209]. On the contrary, previous works had described this normal detachment fault zone as a thrust fault, along which the Kerdylia unit was tectonically placed on the Pangaion unit [202,203,239,240].
At the boundary between the Vertiskos and Kerdylia units crop out tectonically metamorphic mafic and ultramafic rocks of oceanic lithosphere origin, named the Volvi ophiolite complex (Figures 3, 4, 16; [241,242]). Recently, works dated the protolith of the Volvi ophiolite complex as Mid-Late Jurassic age, originating in the Axios/Vardar ocean basin and emplaced in its current position during the Late Cretaceous-Paleocene, following the Axios/Vardar ocean closure and subduction under the European margin [17,24]. Nevertheless, [22,23]suggests an Early Jurassic emplacement of the Volvi ophiolite complex sandwiched between the upper Vertiskos and the lower Kerdylia units and originated in a Volvi-East Rhodope ocean basin. In this scenario, the Kerdylia unit is considered equivalent to the lowermost Pangaion Rhodope unit. A question arises here because the Volvi ophiolite complex protolith has been dated as Mid-Late Jurassic ages. It is impossible that it was emplaced during the Early Jurassic. Furthermore, [200,243] interpret this tectonic contact between Kerdylia and Vertiskos units as a secondary reactivated normal detachment fault zone of the Eocene-Oligocene age along which the overlain Vertiskos unit detached westwards, causing the exhumation of the lower Serbo-Macedonian Kerdylia unit (Figures 3, 4, 16).
According to most recent works (e.g., [17,24,244], the Kerdylia unit is regarded as equivalent to the middle Rhodope Sidironero unit, both forming a single tectonic nappe above the Rhodope`s lowermost Pangaion unit, named the Middle Alochthon while the Pangaion unit as the Lower Allochthon. It is concluded by the recognition of the same structural setting and tectonostratigraphy, as well as the same Eocene-Oligocene cooling/exhumation path for both Sidironero and Kerdylia units (Figures 3, 4, 16; [17,18,24,114,207,225,244]). Moreover, the dating of granitoids of the Kerdylia unit reveals ages of the Late Jurassic and also of the Permo-Carboniferous [245,246], again the same as it is recorded for the Sidironero unit [231]. Nevertheless, the exact ages of high-pressure metamorphic rocks that were recognized in the Serbo-Macedonian Kerdylia unit, remain under debate. A general Mesozoic age was proposed by [211].
  • II. Vertiskos unit
The Vertiskos unit mainly comprises amphibolites, schists, ortho- and paragneisses and in places thin marble layers of a Paleozoic-age protolith [228,247]. Furthermore, abundant migmatite rocks also occur in the lithostratigraphic composition of the Vertiskos unit. Their age remains today under debate, ranging from the Paleozoic to the Tertiary [238,248,249]. A-type leucocratic granitoids (e.g., Arnea granite) intrusions of Early Triassic age (240-220 Ma) are recognized within the Paleozoic rock sequence. They were intruded upon during the continental rifting of the Pangea super-continent (Figure 16; [133,134,246]. Moreover, Neotethyan ophiolite bodies are also tectonically intercalated as imbricated sheets within the Vertiskos basement gneissic and schist rocks (Figures 3, 16; [4,42,238,250]). The ophiolites were initially obducted NE-wards during the Middle-Late Jurassic on the Serbo-Macedonian Vertiskos unit, which formed the European continental margin at the eastern part of the Neotethyan Meliata/Maliac-Axios/Vardar ocean [17,24,42,180,250].
In the Vertiskos unit, similar to the Rhodope Sidironero and Kimi units, high-pressure to ultra-high-pressure diamond-bearing parageneses have been recorded. Their age once again remains under debate. It ranges from Paleozoic to Mesozoic (e.g., [211,248,251]). Moreover, amphibolite facies metamorphic conditions of the Late Jurassic-Early Cretaceous, possibly related to extension and crustal uplift, have been recognized for the Vertiskos unit. It was followed by retrograde greenschist facies Early Cretaceous (Barremian-Aptian) metamorphism related to compressional tectonics (Figure 17b; [198,242,252]). Remnants of Variscan age metamorphism are also recorded [228,238,252,253]. The cooling/exhumation path of the Vertiskos unit has been dated as Early Paleocene-Eocene, while AFT analyses give cooling ages of ~43 Ma (Figures 4, 17b; [225,254]).
Structurally, the Vertiskos unit occupies the same tectonic level as the Rhodope Kimi unit, lying above the Kerdylia unit and shares the same lithostratigraphic composition with the Kimi unit. Additionally, it records a similar tectono-metamorphic evolution as well as cooling/exhumation history to the Kimi unit. These ages are Early Paleocene-Eocene for the Vertiskos unit and Paleocene-Eocene for the Kimi unit. Furthermore, the AFT cooling ages for the Kimi unit reach about 35 Ma [199,225]. Therefore, both units could be regarded as parts of the same tectonic nappe, named “Upper allochthon” according to the subdivision of [17,24,244] tectonically lying above the single Sidironero-Kerdylia nappe (“Middle allochthon”) and which tectonically overlaps the lowermost Pangaion unit (“Lower allochthon”). These nappes column form the Serbo-Macedonian/Rhodope metamorphic nappes stack (Figures 3, 4, 5, 16; [7,24,200,225]). Nevertheless, the somewhat older Early Paleocene-Eocene cooling/exhumation path of the Vertiskos unit, as well as the AFT cooling ages (~43 Ma), show that it was at least one level higher than the Kimi unit or a possibly higher tectonic nappe [18]
The Vertiskos unit is bordered to the West by the Axios/Vardar zone, as earlier in detail described and the Circum-Rhodope belt (CRB). The latter is an Alpine volcano-sedimentary sequence composed of Triassic-Jurassic pelagic and neritic sediments distal and proximal to the continental margin. They include platform carbonate sequences, shales and flysch-type rocks, tectonically intercalated with Triassic bimodal-type volcanic products and acid-to-intermediate island arc ensimatic volcanic rocks formed progressively during an Early-Middle Jurassic intra-oceanic subduction progress in the Neotethyan ocean (Meliata/Maliac-Axios/Vardar ocean). Moreover, an NW-SE striking tectonic slice of pre-Alpine gneisses and schists belonging possibly to the Serbo-Macedonian Vertiskos unit, the so-called Stip-Axios massif, is also tectonically incorporated in between the Mesozoic-Alpine geological formations of the Circum-Rhodope belt (Figures 2, 3, 4, 13, 14; [68,135,237,255,256,257]). Equivalent units to the Circum-Rhodope belt are the Alexandroupolis-Maronia unit in northeastern Greece, as well as the allochthonous Strandja and Mandrica units in Bulgaria (Figures 2, 3; [17,24]. The Circum-Rhodope belt shows a blueschist facies metamorphism of probable Jurassic age, overprinted by a greenschist facies metamorphic event during the late Early Cretaceous [251,256,257,258].
[68,237,256] interpreted the Circum-Rhodope belt as the Alpine, Triassic-Jurassic sedimentary cover of the continental margin of the Vertiskos unit (western European margin) towards the Axios/Vardar ocean basin. According to more recent works [17,24,179,180], the Circum-Rhodope belt and its equivalent units were initially tectonically placed over the Vertiskos margin together with the Neotethyan Axio/Vardar ophiolites with a main NE-ward emplacement direction during the Mid-Late Jurassic in the course of an arc-continent type collision and forming the uppermost tectonic nappe or the “Uppermost allochthon” of the Serbo-Macedonian/Rhodope nappes pile. Through a younger tectonic event during the Late Cretaceous-Paleogene, the initial Mid-Late Jurassic NE-wards stacking direction was reversed to SW-ward thrusting, causing intense imbrication of the CRB unit and the Vertiskos unit. Finally, along an NW-SE striking dextral transpression fault zone of Oligocene-Miocene age the Vertiskos Unit emplaced on the CRB (Figures 3, 4, 10, 13, 14, 16; [6,7,198,256,259]).
Figure 17. a. and b. P/T/t Alpine tectono-metamorphic path and exhumation history of the (a) Rhodope Pangaion and Sidironero units and (b) Serbo-Macedonian Vertiskos unit, AFT=apatite fission-track ages [24,51,59,195,198,225,242]. c. Meso- and microscale compositional features and architecture of the deformational events within the Rhodope and Serbo-Macedonian massifs. Shear sense criteria along X-Z sections [30,198]: I, II. Fossiliferous Triassic-Jurassic carbonate cover of the Serbo-Macedonian continental margin (European margin). Gastropod (I) and ammonite (II) fossils are distinguished; III. D2 isoclinal, recumbent fold of the ?Late Jurassic-Early Cretaceous sedimentary series on the top of the Mid-Late Jurassic obducted ophiolites at the western European margin (Serbo-Macedonian massif). S2 foliation is developed parallel to the fold`s axial plane. (Axios/Vardar zone, Paionia subzone); IV, V. Asymmetric boudins and σ-clasts of leucosoms in Paleocene-Eocene migmatites of the Sidironero unit (Rhodope massif). A top-to-the-SW sense of movement is clearly recognized; VI. S-C-C` fabric of Paleocene-Eocene age in orthogneisses (possible Carboniferous age of the Granite protolith) of the Rhodope Sidironero unit. Sense of shear top-to-the-SW; VII. Sheath-folds, reoriented parallel to the dominant SW-NE trending stretching lineation (X-axis of the strain ellipsoid) in the Pangaion unit`s marbles; VIII. σ- and δ-type feldspar clasts within augen gneiss of the Serbo-Macedonian Vertiskos unit at Kerkini Mt. The sense of shear is top-to-East (D2 event; Barremian-Aptian); IX. Pervasive shear bands and asymmetric quartzitic boudins in schist-gneisses of the Serbo-Macedonian Vertiskos unit. The sense of shear is top-to-NE; X, XI, XII. Shear bands, σ-feldspar clasts and mica “fish” from mica-gneisses of the Serbo-Macedonian Vertiskos unit. Sense of shear clearly top-to-ENE/NE, related to the Paleocene-Eocene D4 event and the exhumation of the Vertiskos unit. The mica notes the S2 foliation but is also reoriented along the D4 shear bands. Microscopic pictures: X, XI, XII with one Nicol (detailed description of the deformational events in chapter “Architecture of deformation and structural evolution”).
Figure 17. a. and b. P/T/t Alpine tectono-metamorphic path and exhumation history of the (a) Rhodope Pangaion and Sidironero units and (b) Serbo-Macedonian Vertiskos unit, AFT=apatite fission-track ages [24,51,59,195,198,225,242]. c. Meso- and microscale compositional features and architecture of the deformational events within the Rhodope and Serbo-Macedonian massifs. Shear sense criteria along X-Z sections [30,198]: I, II. Fossiliferous Triassic-Jurassic carbonate cover of the Serbo-Macedonian continental margin (European margin). Gastropod (I) and ammonite (II) fossils are distinguished; III. D2 isoclinal, recumbent fold of the ?Late Jurassic-Early Cretaceous sedimentary series on the top of the Mid-Late Jurassic obducted ophiolites at the western European margin (Serbo-Macedonian massif). S2 foliation is developed parallel to the fold`s axial plane. (Axios/Vardar zone, Paionia subzone); IV, V. Asymmetric boudins and σ-clasts of leucosoms in Paleocene-Eocene migmatites of the Sidironero unit (Rhodope massif). A top-to-the-SW sense of movement is clearly recognized; VI. S-C-C` fabric of Paleocene-Eocene age in orthogneisses (possible Carboniferous age of the Granite protolith) of the Rhodope Sidironero unit. Sense of shear top-to-the-SW; VII. Sheath-folds, reoriented parallel to the dominant SW-NE trending stretching lineation (X-axis of the strain ellipsoid) in the Pangaion unit`s marbles; VIII. σ- and δ-type feldspar clasts within augen gneiss of the Serbo-Macedonian Vertiskos unit at Kerkini Mt. The sense of shear is top-to-East (D2 event; Barremian-Aptian); IX. Pervasive shear bands and asymmetric quartzitic boudins in schist-gneisses of the Serbo-Macedonian Vertiskos unit. The sense of shear is top-to-NE; X, XI, XII. Shear bands, σ-feldspar clasts and mica “fish” from mica-gneisses of the Serbo-Macedonian Vertiskos unit. Sense of shear clearly top-to-ENE/NE, related to the Paleocene-Eocene D4 event and the exhumation of the Vertiskos unit. The mica notes the S2 foliation but is also reoriented along the D4 shear bands. Microscopic pictures: X, XI, XII with one Nicol (detailed description of the deformational events in chapter “Architecture of deformation and structural evolution”).
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3. Alpine magmatic and depositional processes

For a better understanding and explanation of the Alpine structural-geodynamic evolution of the Hellenides, we consider it important, before the analysis of their structural architecture and kinematics of deformation, to provide here a summary, first (I) of the magmatic activity that affected the Hellenides from the Upper Paleozoic to the Neogene-Quaternary and second (II) of the sedimentary processes recorded in the Hellenides from the Permo-Triassic to the Neogene-Quartenary. Moreover, magmatic and sedimentary processes are very important for dating the several deformational events that affected the Hellenides.

3.1. Hercynian to Neogene-Quaternary magmatism

Initially, Upper Paleozoic (~300 Ma) calc-alkaline granitoid intrusions within the Paleozoic basement units of the Internal Hellenides, related to subduction processes and the Paleotethys closure, have been widely described by several authors (Figures 4, 5, 13, 18; [3,132,133,171,172,260,261]. Early Triassic (~240 Ma) A-type granitoid intrusions and bimodal-type volcanic rocks usually intercalated with clastic and shallow-water carbonate sediments forming a Permo-Triassic volcano-sedimentary series in the Internal Hellenides have been interpreted as magmatism related to the initial Permo-Triassic continental rifting of the Pangea and the Neotethys ocean opening (Figures 3, 4, 16, 18; [130,133,134,262]). Mid-late Middle Jurassic arc-related volcanic products and magmatic activity with granitoid intrusions have been recorded in the Axios/Vardar zone and the Serbo-Macedonian/Rhodope metamorphic province. They intrude the ophiolites and the continental basement rocks and are related to the Meliata/Maliac-Axios/Vardar intra-oceanic subduction processes and the progressively evolved structural stages (Figures 3, 4, 11; [24,41,135,142,164,171,172,263]). Late Cretaceous calc-alkaline arc-type magmatic activity is described for the European margin in the Strandja/Sredna-Gora massif due to the NE-ward subduction of the remnants of the Axios/Vardar oceanic lithosphere under Europe.
This Late Cretaceous magmatic activity migrated to the Southwest, as shown by the SW-ward younger granite intrusion ages from 80 to 70-60 Ma, along the European margin. This magmatic activity was possibly related to a SW-ward rollback of the subduction zone [59,264,265,266,267].
Additionally, Paleocene to Miocene magmatic activity and Tertiary migmatization have been mainly recognized in the Serbo-Macedonian/Rhodope and Cyclades metamorphic provinces, related to extensional regime and/or mantel delamination processes (Figures 3, 4, 16, 19; e.g., [7,17,89,95,197,217,219,220,268]). Finally, Neogene to Quaternary volcanic products are recorded in the broader internal Hellenides zones until the active Hellenic volcanic arc, showing from the older to the younger products an S-SW-ward propagation (Figures 2, 20; [269,270,271,272,273,274].

3.2. Sedimentary processes

Here we briefly describe the main Alpine sedimentary processes of the Hellenides, from the Permo-Triassic to the Neogene-Quaternary. From the older formations to the younger ones, they are: I. The Permo-Triassic volcano-sedimentary sequence with the Triassic-Jurassic neritic and pelagic sediments, proximal and distal of the continental margins (Inernal Hellenides), II. The Mid-Late Jurassic ophiolitic mélanges, III. The Late Jurassic neritic carbonate sediments, IV. The ?late Late Jurassic-Early Cretaceous turbidites and mass-flow deposits, V. The Late Cretaceous shallow-water carbonates and the Paleocene flysch of Internal Hellenides and VI. The Paleocene-Eocene to Oligocene-Miocene flysch sediments of External Hellenides and their Mesozoic basement; the Eocene to Early Miocene turbiditic deposits of the Meso-Hellenic trough and the Thrace basin. Finally, we should also refer to the younger, post-orogenic Hellenides sedimentary series of Neogene-Quaternary age.

3.2.1. Triassic-Jurassic sediments

The Triassic-Jurassic sediments of Internal Hellenides were deposited on the passive continental margins of the Apulia plate (including the Pelagonia) and Europe plate (including the Vertiskos-Kimi unit) in the western and eastern of the Meliata/Maliac-Axios/Vardar ocean basin, respectively, during its opening. These sediments were deposited under an extensional tectonic regime and before the onset of the Alpine deformational history, containing successive nappe stack processes and subsequently extensional events associated with tectonic nappes denudation and crustal uplift [3,6,19,21,130,257,275]. The sedimentary sequences follow a passive continental margin`s typical proximal and distal sedimentation evolution of [137,275]. From the continent to the ocean, they were: neritic platform carbonate sediments, in some places dolomitic; pelagic sediments of the Hallstatt facies; flyshoid sediments of the continental slope of the Meliata facies; and deep sea sediments of the ocean (shales, radiolarites and clay sediments).
Figure 18. Schematic crustal-scale transects showing the geotectonic setting of the Hercynian granitoid intrusions (~300 Ma) into the Pelagonian continental margin and the Permo-Triassic magmatic activity (240 Ma: A-type granitoid intrusions and bimodal volcanism) during the initial stages of the Neotethys opening (modified after [133]). Triassic-Jurassic sedimentation processes take place at both the Pelagonian (Apulia) and Serbo-Macedonian (Europe) passive continental margins. Sedimentation was broken down by the Middle-Late Jurassic Neotethyan ophiolites obduction [6,17,24,142,198].
Figure 18. Schematic crustal-scale transects showing the geotectonic setting of the Hercynian granitoid intrusions (~300 Ma) into the Pelagonian continental margin and the Permo-Triassic magmatic activity (240 Ma: A-type granitoid intrusions and bimodal volcanism) during the initial stages of the Neotethys opening (modified after [133]). Triassic-Jurassic sedimentation processes take place at both the Pelagonian (Apulia) and Serbo-Macedonian (Europe) passive continental margins. Sedimentation was broken down by the Middle-Late Jurassic Neotethyan ophiolites obduction [6,17,24,142,198].
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From the described Triassic-Jurassic sedimentary sequences, the outer shelf sequences are still poorly known because they were strongly reworked during the evolution of the subsequent Jurassic and younger tectonic history. Today, they are detected only in the form of resedimented pebbles or blocks in melanges sequences due to their active participation in the imbrication regime during the Jurassic obduction and the far-traveled advance of the Neo-Tethyan ophiolites on the continental margins [13,30,136,137,275,276,277].
The neritic Triassic-Jurassic sediments rest on top of a Late Permo-Early Triassic volcano-sedimentary series composed of intercalation of clastic sediments, thin carbonate layers and bimodal volcanic products. The Permo-Triassic series evolved, as already mentioned, during the initial break of the Pangea super-continent and the onset of the opening of the Neotethyan ocean [3,130,262].

3.2.2. The Mid-Late Jurassic ophiolitic mélanges

These Mid-Late Jurassic ophiolitic mélange successions, reaching in places several hundred metres thick, are characterized by folds and thrust fault deformation as well as intensive shearing (Figures 21, 22; [20,21,276,278]). They form “block-in-matrix” complexes with a complicated compositional structure composed of up to kilometre-sized exotic blocks (Olistholiths) of Triassic-Jurassic pelagic and shallow-water carbonates sediments, radiolarites, basalts, serpentinites, pillowed and massive lavas and gabbroic rocks, as well as amphibolites from the amphibolite metamorphic sole, which are interbedded in between radiolarites, radiolarian cherts, shales and siliceous shales, dated Bathonian to Oxfordian ages. Furthermore, redeposited fine-grained carbonates and quartzose turbidites are intercalated in places within Mid-Late Jurassic deep-water successions composed of radiolarites, shales and volcanoclastics. Medium to coarsely basic-intermediate sills intrude in places in these lithostratigraphic, polymictic tectono-sedimentary sequences (Figure 22; [13,137,276,277,278,279].
Figure 19. The Tertiary until today SW-ward migration of the dynamic peer compression vs. extension, responsible for the configuration of the Hellenic orogen and its structural architecture, is illustrated (HP=high pressure). The associated Tertiary P-T metamorphic paths of the several high-pressure belts in several regions of the Hellenides are also shown [5,198]).
Figure 19. The Tertiary until today SW-ward migration of the dynamic peer compression vs. extension, responsible for the configuration of the Hellenic orogen and its structural architecture, is illustrated (HP=high pressure). The associated Tertiary P-T metamorphic paths of the several high-pressure belts in several regions of the Hellenides are also shown [5,198]).
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Very well structurally maintained and representative Mid-Late Jurassic ophiolitic mélanges sequences are the Avdella and Vourinos mélanges in the Pindos and Vourinos mountainous regions, respectively [115,267,278,280]. In Orthrys Mountain, as well as in the Axios/Vardar zone at its both western and eastern marginal parts, similar ophiolitic mélange formations are also described [21,27,37,42,181,281,282]. They are sandwiched on the one hand between the External Hellenides Pindos flysch at the bottom and the detached ophiolite belt at the top and on the other hand between the Pelagonian (Vourinos Mountain) or Serbo-Macedonian Triassic-Jurassic carbonate platform cover at the bottom and the obducted ophiolites at the top (Figures 4, 11, 21, 22). The internal deformation of the mélange formations shows a main sense of movement ranging from top-to-the-NW to the-SW for the successions on top of the Pelagonian nappe at the western of the Axios/Vardar marginal edge and top-to-the-NE for the successions on top of the Serbo-Macedonian at the eastern of the Axios/Vardar marginal edge, respectively. Sense of movement is recognized by shear sense indicators such as σ- and δ-clasts, shear bands, asymmetric boudins and thrust vergence (Figure 22). It is interpreted as the kinematics, related to the initial ophiolites and mélanges nappe thrusting and their final emplacement on the Pelagonian and Serbo-Macedonian continental margins during the Mid-Late Jurassic [6,13,30,42,119,120,137,148,179,180,183,275]. The emplacement on the Pindos flysch, together with the overlain ophiolites, clearly westwardly directed, belongs to the younger.
Figure 20. The SSW-ward migration of the Late Cretaceous to present-day arc-type magmatic activity in the Hellenic arc is shown. The main tectonic units of southeastern Europe and the eastern Mediterranean region are aso indicated. Abbreviations as in Figure 3.
Figure 20. The SSW-ward migration of the Late Cretaceous to present-day arc-type magmatic activity in the Hellenic arc is shown. The main tectonic units of southeastern Europe and the eastern Mediterranean region are aso indicated. Abbreviations as in Figure 3.
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Tertiary tectonics of the Hellenides, related either to compressional or extensional tectonics [19,276,278,283,284,285]. During this Tertiary tectonics, the ophiolite nappe together with the underlain ophiolitic mélanges were simply detached on the External Hellendes Pindos flysch without any important effect or reworking on the older Jurassic inherited nappe thrusting structures (Figures 3, 4, 21; [119,120,125,183,283,284]).
These Mid-Late Jurassic ophiolitic mélanges were interpreted as a synorogenic, tectono-sedimentary sequence, deposited in progressively formed Mid-Late Jurassic basins at the front of the advanced ophiolites thrust sheets, following the Early-Middle Jurassic intra-oceanic subduction and partly destruction of the Neotethyan ocean floor (Meliata-Axios/Vardar ocean; [6,13,119,120,137,148,183,275]). Amphibolite metamorphic sole always of 180-170Ma age [158,286,287], tectonically incorporated in between all ophiolite belts, show clearly this late Early-Middle Jurassic intra-oceanic subduction event, only in a single ocean basin and this should be the Neotethyan Meliata/Maliac-Axios/Vardar ocean between the Apulian and European plates (e.g., [11,13,17,24,275]).
Ophiolitic mélanges of similar composition and Mid-Late Jurassic age are also described by several authors in the northern continuation of the Hellenides ophiolites in the Albanian Mirdita ophiolite belt, as well as the Dinarides and eastern Alpen (e.g., [13,19,137,159,275]. These mélange formations were interpreted as radiolarite-ophiolitic wild flysch [13] and these correspond to similar sequences known from the eastern Alpen region (Hallstatt mélanges; [288]).
Figure 21. Geological-structural map and representative cross-section of the Pindos ophiolites and the tectonically underlying Middle-Late Jurassic Avdella ophiolitic mélange that are tectonically emplaced on the Paleocene-Eocene Pindos flysch. Outcrops of the sedimentary formations of the western molassic Mesohellenic trough and its tectonic boundary with the basement rocks (ophiolites, Late Cretaceous limestones and Pindos flysch) are also shown (based on [64,65,276,278]). Abbreviations as in Figure 3.
Figure 21. Geological-structural map and representative cross-section of the Pindos ophiolites and the tectonically underlying Middle-Late Jurassic Avdella ophiolitic mélange that are tectonically emplaced on the Paleocene-Eocene Pindos flysch. Outcrops of the sedimentary formations of the western molassic Mesohellenic trough and its tectonic boundary with the basement rocks (ophiolites, Late Cretaceous limestones and Pindos flysch) are also shown (based on [64,65,276,278]). Abbreviations as in Figure 3.
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3.2.3. The Late Jurassic sedimentary series

These sedimentary series comprise shallow-water fossils-rich platform limestones
of the Kimmeridgian-Tithonian age, unconformably deposited on the ophiolites directly after their emplacement on the continental margins of the Pelagonian and Serbo-Macedonian (Figures 3, 4, 5, 15, 23 VIII, IX; [13,19,21,147,148,277]). Furthermore, oolithic limesto-ne layers constitute, also in some places (e.g., the Koziakas unit), a common compositional component of the lithostratigraphic succession. This Late Jurassic shallow-water carbonate formation today remains only as several small rests on its basis (Figure 23 VIII) because its greater mass is eroded or, in some places totally eroded. Its age and existence are proven by the dating of its eroded compositional material contained as carbonate clasts and blocks in mass-flow and clastic formations, which have been dated as ?late Late Jurassic-Early Cretaceous age and are described just below [6,137,147,148,149,150,275,277].
Figure 22. I to XII. Field photos of structural and compositional features of the ophiolite belt and the ophiolitic mélanges, as well as the deep sedimentary sequences of the ?Sub-Pelagonian zone that underlie the ophiolites and ophiolitic mélanges (modified after Kilias et al. 2016). The sense of shear is indicated by arrows and the respective tectonic events are shown in each photo. The Early-Middle Jurassic amphibolite metamorphic sole, formed during the Early-Middle Jurassic Neotethyan intra-oceanic subduction between the overriding hot lithospheric mantel (the serpentinites) and the ophiolitic mélanges, is also shown (X, XI). The (I, II, III, XII) photos refer to the ophiolites of the Axios/Vardar zone along the eastern Pelagonian margin and the (IV, V, VI, VII, VIII, IX) photos refer to the Pindos ophiolite belt and ophilolitic mélanges, as well as the ?Sup-Pelagonian pelagic sediments along the western Pelagonian margin. Abbreviations as in Figure 3.
Figure 22. I to XII. Field photos of structural and compositional features of the ophiolite belt and the ophiolitic mélanges, as well as the deep sedimentary sequences of the ?Sub-Pelagonian zone that underlie the ophiolites and ophiolitic mélanges (modified after Kilias et al. 2016). The sense of shear is indicated by arrows and the respective tectonic events are shown in each photo. The Early-Middle Jurassic amphibolite metamorphic sole, formed during the Early-Middle Jurassic Neotethyan intra-oceanic subduction between the overriding hot lithospheric mantel (the serpentinites) and the ophiolitic mélanges, is also shown (X, XI). The (I, II, III, XII) photos refer to the ophiolites of the Axios/Vardar zone along the eastern Pelagonian margin and the (IV, V, VI, VII, VIII, IX) photos refer to the Pindos ophiolite belt and ophilolitic mélanges, as well as the ?Sup-Pelagonian pelagic sediments along the western Pelagonian margin. Abbreviations as in Figure 3.
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3.2.4. The ?late Late Jurassic-Early Cretaceous mass flow and clastic sedimentary series

These sedimentary series also rest unconformably, mainly directly on top of the ophiolitic belts. (Figure 3, 4, 5, 10, 11, 23). These sediments form very complicated reef slope rock units but are very important for the explanation of the structural-geotectonic evolution of the Internal Hellenides. They form polymictic mass-flow deposits containing a large number of carbonate components from the former described and intensively eroded Late Jurassic shallow-water carbonate platform, as well as from the underlain ophiolites and basement rocks (e.g., Pelagonian or Serbo-Macedonian Triassic-Jurassic carbonate platform). Radiolaritic and flyschoid clasts also occur as components in the mass flow. As components of the eroded Late Jurassic carbonate platform dominate resedimented bio- oclasts of reefal limestones, shells, stromatoporoids, corals, brachiopods, foraminifera, crinoids and sponges. Some fragments are coated and encrusted (Figures 23, 24; [148,150]).
Figure 23. Field photos of the Late Jurassic and ?Late Jurassic-Early Cretaceous carbonate sequences on the top of the Late Jurassic obducted ophiolites on the Pelagonian and Serbo-Macedonian continental margins: I. The deposition of the ?Late Jurassic-Early Cretaceous carbonate neritic and clastic sediments over the Axios/Vardar Paionia ophiolite belt at the Serbo-Macedonian margin; II, III. The carbonate clasts and the thin carbonate layer at the base of the ?Late Jurassic-Early Cretaceous sedimentary sequence is rich in fossils representative of a Late Jurassic fauna collection. Axios/Vardar zone, Paionia Subzone, Serbo-Macedonian margin; IV, V, VI, VII. The Early Cretaceous clastic and carbonate semi-pelagic sediments on top of the Vourinos ophiolites complex at the western Pelagonian flank. Clastic sediments with small belemnite fragments (IV) and calpionellides bearing turbidity layers (V) are distinguished. Furthermore, the Early Cretaceous compressional D2 event related to subisoclinal, asymmetrical folds is shown (VI, VII); VIII, IX. Is shown the contact between the Late Jurassic shallow-water limestones and the Zygosti ophiolites, which form the W-ward continuation of the Axios/Vardar ophiolites (VIII). Fauna from the rich fossiliferous neritic Late Jurassic limestones at the Zygosti stream, covering the obducted ophiolites belt on the Pelagonian continent (IX) (based on [30,149,150]).
Figure 23. Field photos of the Late Jurassic and ?Late Jurassic-Early Cretaceous carbonate sequences on the top of the Late Jurassic obducted ophiolites on the Pelagonian and Serbo-Macedonian continental margins: I. The deposition of the ?Late Jurassic-Early Cretaceous carbonate neritic and clastic sediments over the Axios/Vardar Paionia ophiolite belt at the Serbo-Macedonian margin; II, III. The carbonate clasts and the thin carbonate layer at the base of the ?Late Jurassic-Early Cretaceous sedimentary sequence is rich in fossils representative of a Late Jurassic fauna collection. Axios/Vardar zone, Paionia Subzone, Serbo-Macedonian margin; IV, V, VI, VII. The Early Cretaceous clastic and carbonate semi-pelagic sediments on top of the Vourinos ophiolites complex at the western Pelagonian flank. Clastic sediments with small belemnite fragments (IV) and calpionellides bearing turbidity layers (V) are distinguished. Furthermore, the Early Cretaceous compressional D2 event related to subisoclinal, asymmetrical folds is shown (VI, VII); VIII, IX. Is shown the contact between the Late Jurassic shallow-water limestones and the Zygosti ophiolites, which form the W-ward continuation of the Axios/Vardar ophiolites (VIII). Fauna from the rich fossiliferous neritic Late Jurassic limestones at the Zygosti stream, covering the obducted ophiolites belt on the Pelagonian continent (IX) (based on [30,149,150]).
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Calpionellids bearing pelagic limestones, which date as Early Berriasian [153,289] followed by calcareous-siliciclastic turbidites of Late Berriasian age [290] overlain the mass-flows sequence (Figure 23). The whole succession, of mass flow deposits with the Calpionellid limestones was dated as Early Cretaceous (Berriasian-Valanginian; [13,147,148,149,150,277]). A late Late Jurassic (late Tithonian) age remains questionable. Moreover, in some places the stratigraphic succession through shallow-water carbonate rocks and flysch-like deposits reaches until the Aptian [13,21,137,143,277,291].
Figure 24. Components of the ?Late Jurassic-Early Cretaceous polymictic mass flows and clastic sedimentary sequence: I. Rudstone with different clasts and Labyrinthina mirabilis Weynschenk (width: 0.5 cm); II. A fragment of Griphoporella jurassica (Endo) (width: 0.25 cm); III. Neoteutloporella socialis (Praturlon) (width: 0.5 cm); IV. Boundstone with Thaumatoporella sp (width: 0.5 cm); V. Boundstone with Perturbatacrusta leini Schlagintweit and Gawlick (above) and Labes atramentosa Eliasova (below) (width: 0.5 cm). Components from the ?Late Jurassic-Early Cretaceous turbiditic sequence of sandstones with intercalated coarse-grained mass flows; VI. Anchispirocyclina lusitanica (Egger) (width: 0.5 cm); VII. Grainstone with different clasts and Salpingoporella pygmaea (Gümbel) (width: 0.5 cm). Components from the polymictic conglomerate in the ?Late Jurassic-Early Cretaceous sedimentary sequence; VIII. Metamorphosed shallow-water clasts of most probably Middle Triassic age containing foraminifera (width: 0.5 cm). Microfossils from the shallow-water ?Late Jurassic-Early Cretaeous sedimentary sequence; IX. Griphoporella cretacea (Dragastan) (width: 0.5 cm); X. Boundstone with Suppiluliumaella aff. methana Dragastan and Richter and debris of Selliporella neocomiensis (Radoicic) (width: 0.5 cm); XI. Linoporella aff. capriotica (Oppenheim) (width: 0.25 cm); XII. Furcoporella? vasilijesimici Radoicic (width: 0.25 cm). [149,150].
Figure 24. Components of the ?Late Jurassic-Early Cretaceous polymictic mass flows and clastic sedimentary sequence: I. Rudstone with different clasts and Labyrinthina mirabilis Weynschenk (width: 0.5 cm); II. A fragment of Griphoporella jurassica (Endo) (width: 0.25 cm); III. Neoteutloporella socialis (Praturlon) (width: 0.5 cm); IV. Boundstone with Thaumatoporella sp (width: 0.5 cm); V. Boundstone with Perturbatacrusta leini Schlagintweit and Gawlick (above) and Labes atramentosa Eliasova (below) (width: 0.5 cm). Components from the ?Late Jurassic-Early Cretaceous turbiditic sequence of sandstones with intercalated coarse-grained mass flows; VI. Anchispirocyclina lusitanica (Egger) (width: 0.5 cm); VII. Grainstone with different clasts and Salpingoporella pygmaea (Gümbel) (width: 0.5 cm). Components from the polymictic conglomerate in the ?Late Jurassic-Early Cretaceous sedimentary sequence; VIII. Metamorphosed shallow-water clasts of most probably Middle Triassic age containing foraminifera (width: 0.5 cm). Microfossils from the shallow-water ?Late Jurassic-Early Cretaeous sedimentary sequence; IX. Griphoporella cretacea (Dragastan) (width: 0.5 cm); X. Boundstone with Suppiluliumaella aff. methana Dragastan and Richter and debris of Selliporella neocomiensis (Radoicic) (width: 0.5 cm); XI. Linoporella aff. capriotica (Oppenheim) (width: 0.25 cm); XII. Furcoporella? vasilijesimici Radoicic (width: 0.25 cm). [149,150].
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Where these both Late Jurassic and Early Cretaceous sedimentary formations were mapped on top of the obducted ophiolites in several localities of the Alpine orogenic belt from the eastern Alps, Dinarides Albanides until the Hellenides (Figures 1, 3, 4, 10, 15; e.g., western and eastern boundaries edges of the Pelagonian nappe, Axios/Vardar zone and Serbo-Macedonian province), they show an identical stratigraphical succession, similar sedimentary features and components composition and were dated with about the same stratigraphic age: I. Late Jurassic (Kimmeridgian-Tithonian) for the first, intensively eroded shallow-water carbonate platform formation and II. ?late Late Jurassic-Early Cretaceous for the second one. Additionally, these strongly eroded Late Jurassic (Kimmeridgian-Tithonian) shallow-water limestones on top of the obducted ophiolitic belts clearly define the upper limit of the ophiolites` emplacement as Callovian-Oxfordian, everywhere in the Alpine orogenic belt in Europe (Alps, Dinarides, Albanides and Hellenides). Moreover, the ophiolite belts in Hellenides, whether they are displayed at the western or eastern Pelagonian parts, even at the Serbo-Macedonian margin, should have the same emplacement age (Figures 2, 3, 4; [13,21,137,149,150,277]).

3.2.5. The Late Cretaceous shallow-water carbonate formation

This formation was deposited unconformably upon all pre-Late Cretaceous geological units and structures of the Internal Hellenides. The deposition began during the Cenomanian. So, we meet it transgressively above the Mid-Late Jurassic obducted ophiolite belts, the Triassic-Jurassic Pelagonian and Serbo-Macedonian platform carbonate covers, the Late Jurassic neritic limestones and the Early Cretaceous mass flow deposits (Figures 3, 4, 5, 11). The formation is composed mainly of neritic carbonate deposits terminated with a late Late Cretaceous-Paleocene flysch sedimentation, known as the flysch of the Internal Hellenides.
A detailed, representative stratigraphic column of the formation, from the bottom to the top, is given in the following: The base of the Late Cretaceous succession forms a few metres of thick conglomerates. They are dominated by clasts of marbles, ophiolites and schists, in some places with a matrix of reworked ophiolitic material or thin layers of terrigenous ophiolitic fine-grained material and shales. The succession is continued with a thin dolomitic layer passing upwards into intercalations of thin- to medium-bedded bioclastic, neritic limestones, mudstones, terrigenous sandstones and redeposited limestones. In places, debris flow material rich in limestone clasts is also observed in between the limestone layers. In the uppermost part, semi-pelagic limestones dominate and the rocks show a deepening of the basin until the final deposition of the Maastrictian-Danian turbiditic deposits. The overall thickness of the Late Cretaceous carbonate succession ranges accordingly, from a few metres to ~2km [21,27,160,161,162,292].

3.2.6. The External Hellenides flysch series and the turbiditic deposits of the Meso-Hellenic trough and the Thrace basin. The post-orogenic Neogene-Quaternary sediments.

The External Hellenides are characterized by a Paleocene to Miocene synorogenic flysch sedimentation. Its age was progressively dated younger W-ward, simultaneously with its W-ward migration. We distinguish from the East to the West: I. The Pindos Paleocene-Eocene flysch; II. The Gavrovo Eocene-Oligocene flysch; and III. The Ionian Oligocene-Miocene flysch. The flysch sediments generally sealed progressively the sedimentary history of the External Hellenides zones, which is characterized by continuous neritic and pelagic sedimentation during the Early Mesozoic to the Tertiary (details described in Chapter 2.1). The flysch sediments were developed during the finale collision of the Apulian origin`s Pelagonian continental fragment with the Apulia plate, closely related to intensive compressional and W-SW-directed folds and thrust faults tectonics (Figures 1, 2, 3, 4; [25,26,136,278]).
Furthermore, very important for the geological history of the Hellenides are the mollasic-type turbiditic syn- to late orogenic, Paleocene to Middle-Late Miocene sedimentary deposits of the Meso-Hellenic trough, defined as a polyhistor strike-slip and piggyback basin, behind the Tertiary External Hellenides thrust belt. Finally, here should also be referred to the Eocene-Oligocene turbidites intercalations with volcanic products of the supra-detachment Thrace basin in the Rhodope province (Figures 1, 2, 3, 4; [7,25,61,62,63,64,65,67]).
Post-orogenic Neogene-Quaternary sedimentary series deposited in intramontagne and other terrestrial sedimentary basins cover, unconformably in many places, all the pre-Alpine and Alpine Hellenides tectonostratigraphic domains. These basins are usually bounded by high-angle normal faults (Figures 1, 2, 3, 4; e.g., [68,69,70,71,72,73,293]).

4. Architecture and kinematics of deformation and the related metamorphic conditions

From the Early-Middle Jurassic to the present, the Hellenides were affected by complicated multiphase stages of deformation and metamorphism during the Alpine orogeny. Deformation started with an Early-Middle Jurassic intra-oceanic subduction followed by the Middle-Late Jurassic ophiolites obduction. The following, younger deformational and metamorphic history, from the Late Jurassic to the present, was divided into six main (D1 to D6) deformational events. (Figures 4, 5, 25, Table 1). The whole Alpine deformational, metamorphic and magmatic history of the Hellenides is described in detail below. In some places in the Paleozoic basement rocks of the Internal Hellenides an older deformational event of Ercynian age is also recognized, however strongly overprinted by the younger Alpine deformation [6,7,30,131,142,199,294].
In a general view, during the Alpine orogeny in the Hellenides, compression, nappe stacking and crustal thickening alternated successively through time with extension and orogenic collapse that led to the exhumation of deep crustal levels, crustal thinning and the formation of tectonic windows and/or metamorphic core complexes. The exhumation structural processes were accordingly evolving from ductile to brittle deformation conditions. Furthermore, an S-SW-ward migration of the dynamic peer compression vs. extension is recognized during the Tertiary Alpine orogenic stages in the Hellenides. Extension and crustal uplift follow compression and nappe stacking or act simultaneously in different sections of the Hellenic orogen (Figures 4, 19, 20; [9,10,32,33,48,89,90,136,156,198,295,296,297]).

4.1. Late Jurassic-Early Cretaceous deformational event (D1) and associated pre-Late Jurassic Alpine structural processes (~180 to 130 Ma)

The D1 event is recognised on the Internal Hellenides, including the Pelagonian nappe, the Sup-Pelagonian zone as defined, the Axios/Vardar zone, the Circum-Rhodope belt and the Serbo-Macedonian massif [3,6,37,142,151,179,180,181,198,242].
The D1 event is dated to Late Jurassic-early Early Cretaceous age (150-130 Ma; [151,298]) and is characterized by a penetrative syn-metamorphic foliation (S1) associated with residual forms of isoclinals recumbent, as well as sheath folds (F1), which deform a pre-existing foliation (S0). The F1-fold axes develop parallel to the mineral stretching lineation (L1), which usually trends NE-SW, but in some places an NW-SE trend is also recognized, for example in the lower Pelagonian segment in Voras Mountain and the Circum-Rhodope belt, as well as in Paleozoic basement slices incorporated in the Axios/Vardar zone (Figures 11b, c, 25; [6,151]). No clear kinematic indicators of the D1 event have been preserved because the D1 structures have been strongly overprinted by the subsequent tectonics. However, in a few places where the D1 kinematics is recognized, a main sense of movement top-to-the-SW or top-to-the-NE may be reconstructed. An NW-ward sense of movement is also recorded, e.g., in the lower Pelagonian segment in northern Greece (Figure 25; [6,41,42,142,179,180,181,198]). Furthermore, an important bulk coaxial component of the D1 deformation is also recognized [6,142,198]. The syn-D1 mineral parageneses, also identified in the L1-stretching lineation, show metamorphic conditions from the greenschist to amphibolite facies, from the structurally higher to the structurally deeper crustal levels, respectively (Figures 11a, b, c, 17b; [6,142,242]).
High-pressure metamorphism in the Internal Hellenides, predating the D1, has been suspected in several works by residuals only of high-pressure metamorphic assemblages and without any clear syn-metamorphic structural evidence due to the strong overprinting by the younger tectonics [6,52,54,142,151,258,298,299]. High-pressure metamorphism is normally related to compression and subduction processes, but in our case, high-pressure metamorphism has been interpreted as the result of tectonic overpressure and continental crustal sinking due to overloading and nappe stacking in an arc-continent collision setting and the ophiolites obduction on the Pelagonian and Serbo-Macedonian/Rhodope continental margins, respectively [6,21,142]. Pre-D1 high-pressure metamorphic assemblages have been dated to early Late Jurassic age (>150 Ma, Figures 11a, b, 17b; [6,151,258,298,299]).
Here, we must also mention the very important formation of the metamorphic sole of the Early-Middle Jurassic age, developed during the Early-Middle intra-oceanic subduction in the Neothyan ocean realm (180-170 Ma, Figure 22; [158,286,287]). This event, in any case, predates the Middle-Late Jurassic ophiolite`s obduction, the high-pressure metamorphism and the D1 event but is closely related to their evolution (Figures 3, 4; [6,17,24,119,120,148,179,180,181,183]). Amphibolite metamorphic sole predates with the same ages the obduction of the ophiolites belt either on the Pelagonian or the Serbo-Macedonian margins, indicating at least its simultaneous evolution at a similar origin`s source and this was the Early-Middle Jurassic intra-oceanic subduction in the Neotethyan Meliata/Maliac-Axios/Vardar ocean basin (Figure 22; [6,13,24,119,120,183,258]). Intra-oceanic subduction is associated with Middle-late Middle Jurassic island arc magmatism now represented mainly as slices in the rock sequences of the Circum-Rhodope belt and the Axios/Vardar zone (Figures 3, 4, 5, 10, 14; [135,142,164,256,257]). Furthermore, the previously described Middle-Late Jurassic ophiolitic mélanges were deposited and intensively imbricated in basins at the front of the advanced ophiolites during their inneroceanic thrusting and subsequently obduction on the Pelagonian and Serbo-Macedonian continental margins. The upper limit of the ophiolites obduction, as previously mentioned, is defined as Callovian-Oxfordian age, much earlier than previously assumed (Figures 15, 23VIII, IX; [13,19,21,147,148,276,277,278,284,300]).
Taking into account: I. The previously described intensively eroded Late Jurassic platform carbonate sediments (Kimmeridgian-Tithonian) on top of the obducted ophiolites; II. The subsequent Early Cretaceous resedimented mass-flow sedimentary series, containing components of the eroded Late Jurassic sediments; and III. the P/T/t metamorphic path during the Middle-Late Jurassic to Early Cretaceous for the Pelagonian and Serbo-Macedonian basements (Figures 11a,b, 17b), it was proposed an uplifting process during the D1 Late Jurassic-Early Cretaceous event for the Pelagonian and Serbo-Macedonian continental margins, respectively [6,119,120,126,127,134]. Uplift took place after the Mid-Late Jurassic Neotethyan Axios/Vardar ophiolites obduction (Callovian-Oxfordian) and the high-pressure metamorphic event and was associated with extension and exhumation of the Pelagonian and Serbo-Macedonian marginal parts [6,127]. This crustal uplift and exhumation could be attributed to buoyancy forces due to the lighter, former submerged continental crust of the obducted ophiolites overload or, additionally, slab break-off of the subducted lithosphere [41,177]. Furthermore, isoclinal, recumbent folds described during the D1 have also been described during synorogenic crustal extension and uplift in the Betic Cordilleras (Spain) by [301,302], as well as in the Alps (Austroalpin domain) by [303,304].

4.2. Late Early Cretaceous deformational event (D2; ~125-115 Ma)

The D1 and the pre-D1 related older structures are strongly overprinted by a younger, contractional, D2 syn-metamorphic deformation, recognized everywhere in the Internal Hellenides zones. The D2 event has strongly affected all pre-late Early Cretaceous (Albian) geological units. They include the Paleozoic basement rocks with their Triassic-Jurassic sedimentary cover and the obducted ophiolite belt, as well as the Early Creta- ceous mass flows and turbiditic sedimentary series. Additionally, the D2 contractional structures are sealed by the Late Cretaceous neritic carbonate deposits (Figures 13, 14, 15). They clearly indicate a late Early Cretaceous age for the D2 event (Figures 3, 4, 11a, b, 13, 14; [6,43,142,154,294]). Furthermore, isotopic dating analyses show similar late Early Cretaceous age deformation in the Pelagonia and Axios/Vardar zone (Figure 11a, b; Barremian-Aptian, 125-115 Ma; [151,298]).
Figure 25. In comparison, the main deformational events recorded in the External and Internal Hellenides from the Early Triassic to the present. Shortening alternated successively with extension. Compressional events are related to nappe stacking, crustal thickening and high-pressure metamorphism, while extensional ones are related to tectonic denudation, low- to high-temperature metamorphism, isostatic rebound, crustal thinning and exhumation of deep crustal levels (see also Figures 27, 28). The main sense of movement for each event, as it was deduced by shear sense indicators and kinematic analysis, is shown in the Schmidt diagrams with arrows (lower hemisphere equal-area projections). (based on [6,30,142]).
Figure 25. In comparison, the main deformational events recorded in the External and Internal Hellenides from the Early Triassic to the present. Shortening alternated successively with extension. Compressional events are related to nappe stacking, crustal thickening and high-pressure metamorphism, while extensional ones are related to tectonic denudation, low- to high-temperature metamorphism, isostatic rebound, crustal thinning and exhumation of deep crustal levels (see also Figures 27, 28). The main sense of movement for each event, as it was deduced by shear sense indicators and kinematic analysis, is shown in the Schmidt diagrams with arrows (lower hemisphere equal-area projections). (based on [6,30,142]).
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D2 is characterized by the evolution of thrust faults and the development of micro- to macro-scale asymmetric, recumbent to overturned, tight to isoclinal folds (F2). They refold both the S1-foliation and the isoclinal F1 folds. A new S2 foliation was developed parallel to the F2 folds axial planes, forming in places a well-recognized crenulation cleavage with the S1 foliation. Usually, due to the intensive rotation of S1 along the S2, the S1 is parallelized to the S2 foliation. As a result, the two foliations coincide and only one foliation is observed (Figures 11c, I, III, VII, XI, 25). The F1 and F2 fold axes are developed mostly parallel to each other, while in some places the F2 folds show even a scattering of their trend, from NW-SE to NE-SW. We attribute this scattering of the F2 folds trend to a progressive rotation towards the stretching direction and the X-strain axis developed during the D2 deformation. The stretching lineation trend is mainly NE-SW, but in some cases, an NW-SE trend is also observed, as is concluded by the development of the syn-D2 metamorphic mineral parageneses on the S2 foliation planes. The sense of shear during the D2 is according to the stretching lineation trend and the shear sense indicators, mainly top-to-the-SW in the Pelagonia and Axios/Vardar zone and top-to-the-NE in the Serbo-Macedonian province. Additionally, an NW-ward sense of movement is also observed, mainly in the lower Pelagonian segment and the Axios/Vardar zone. We consider this geometry of the D2 deformation as the result of a transpressional tectonic regime (Figures 3, 25; [6,17,24,43,154,155,179,180]). M2-metamorphic mineral parageneses show metamorphic conditions in the greenschist facies (Figure 11a, b, c; [6,142,151]). Recent works by [43], [154] mention late Early Cretaceous ages of ca. 117 Ma for migmatites in the deeper Pelagonian levels.

4.3. Late Early Cretaceous-Late Cretaceous deformational event (D3; ~110-80 Ma)

The late Early Cretaceou-Late Cretaceous event is mainly recognized in the Internal Hellenides; in the Pelagonian nappe and its metamorphic basement sheets in the Axios/Vardar zone (e.g., Peternik, Paikon etc. basement rocks), as well as in the Serbo-Macedonian and Rhodope massifs.
In the Pelagonian nappe and the Axios/Vardar zone basement sheets, western edge of the Axios/Vardar ocean, D3 is related to extension and has been dated of late Early to early Late Cretaceous age (110-95 Ma, Figure 11a, b, c; [6,30,151,298]) The D3 extensional structures form discrete, mylonitic shear bands with a well-developed S3-mylonitic foliation dipping mainly towards the NE. The associated L3-stretching lineation also plunges to the NE. The D3 shear bands are usually characterized by the dynamic recrystallization of quartz and the growth of chlorite and sericite minerals. The sense of shear during the D3 was mainly identified down-dip towards NE, although in a few cases an opposite, again down-dip, towards SW sense of shear is also observed, related to SW-ward dipping of the S3-foliation and SW-ward plunge direction of the L3 stretching lineation (Figures 11c, V, VI, 25; [6,151,298]).
In contrast to the described D3 extensional structural setting in the Pelagonian and Axios/Vardar basement rocks, in the Serbo-Macedonian/Rhodope province, Late Cretaceous contraction is revealed by the eclogites occurrences of ca. 80-85 Ma in between the Rhodope units and in detail between the middle Rhodope Sidironero and the upper Rhodope Kimi units. Eclogites are related to ongoing NE-ward subduction processes of the Axios/Vardar oceanic lithosphere remnants under the European continental margin, including the Serbo-Macedonian/Rhodope single Vertiskos-Kimi unit and the Strandja/Sredna-Gora massif in Bulgaria. The subduction processes are also inferred by Late Cretaceous calc-alkaline volcanic-arc magmatism along the southwestern European continental margin (Figures 2, 20; [24,59,117,180,264,266]). Therefore, according to the D3- described structural setting, Late Cretaceous contraction at the eastern-most area of the Internal Hellenides in the Serbo-Macedonian/Rhodope province (European margin) takes place about simultaneously with late Early Cretaceous-early Late Cretaceous extension in the Pelagonia (Apulian margin).

4.4. Early Paleocene to Early Oligocene deformational event (D4; ~65-30 Ma)

The D4 event is also characterized by a very complicated evolutionary history. It is related to ductile up to brittle deformational conditions in the different Hellenides orogenic parts, causing either intense folding, imbrication and nappes stacking during compressional setting or nappes denudation, orogenic collapse and crustal exhumation during extension (Figures 3, 4). The D4 structures are well recognized in the Pindos zone of the External Hellenides, as well as in the Pelagonian nappe pile, the Axios/Vardar zone and the Sebo-Macedonian/Rhodope metamorphic province of the Internal Hellenides (Figures 11c, 26).
Ductile and brittle deformation during D4 takes place simultaneously at the various structural levels of the Hellenic orogenic belt. So that ductile deformation in the Serbo-Macedonian/Rhodope metamorphic province and the lower structural levels of the Pelagonian nappe, including the Paleocene-Eocene Ampelakia-Cyclades high-pressure belt, takes place at the same time as brittle deformation at the upper structural levels of the Pelagonian nappe pile, the Axios/Vardar zone and the structural province of the external parts of the Pindos zone. The brittle deformation is characterized by intense crustal imbrication and thrust faults, as well as asymmetrical kink-folds (Figure 26). Additionally, the related ductile D4 structures are contractional in the middle Rhodope Sidironero unit, the Pelagonian nappe and the Ampelakia-Cyclades high-pressure belt, related to asymmetrical, tight to isoclinal folds and syn-metamorphic S4 schistosity. They today remain residual or destroyed due to the younger tectonics.
Furthermore, during the Paleocene-Eocene, the collision between Europe and Apulian`s Pelagonian segment takes place, following the finale closure of the Axios/Vardar ocean [17,24,179]. Progressively, during this contractional stage of the D4 tectonics, the middle Serbo-Macedonian/Rhodope Sidironero-Kerdylia unit overthrusts the lowermost Rhodope Pangaion unit. About simultaneously, during the Eocene-Oligocene, the Pelagonian nappe and the Paleocene-Eocene Ampelakia-Cyclades high-pressure belt overthrust the Olympos-Ossa carbonate unit of the External Hellenides. On the contrary, the D4 ductile structures are extensional in the uppermost structural nappes of the Serbo-Macedonian/Rhodope metamorphic province, i.e., the Vertiskos-Kimi unit and these structures are ductile shear zones, mainly SW-ward down-dip verging and in some places opposite NE-ward down-dip also verging. Remnants of asymmetrical and isoclinal or sheath folds are also sporadically recognized (Figures 3, 4; [6,7,17,18,24,27,30,37,38,39,47,119,120,142,153,184,196,224,294]).
The D4 deformation, whether ductile or brittle, took place during the Paleocene to Eocene-Oligocene, as is concluded by numerous isotopic dating ages and lithostratigraphic-structural relationships [3,6,7,24,27,30,37,38,39,87,151,160,161].
The residual ductile compressional D4 structures in the lowermost Pelagonian parts, the internal high-pressure belt in Ampelakia-Cyclades and the Rhodope Sidironero-Kerdylia unit are all associated with high-pressure metamorphic conditions (Figures 4, 8a, b, c). On the contrary, the ductile D4 extensional step took place under greenschist to amphibolites facies metamorphic conditions and partly migmatization, as well as granitoid intrusions, associated with the exhumation of the upper Serbo-Macedonian/Rhodope Vertiskos-Kimi unit (Figures 4, 16, 17b, c; [7,39,40,45,47,197,198,199,225]).
A main SW-ward sense of movement is recognized for all ductile or brittle D4 structures, whether they are related to contraction or extension. In some cases, a NE-ward back-direction of sense of movement during the D4 event is also identified in the broader area of the internal Hellenides, the same if the structures are related to contractional or extensional settings (Figures 17c, 25, 26; [6,7,18,27,37,40,142,160,161,198,305]).

4.5. Oligocene-Miocene deformational event (D5; ~30Ma-20 Ma)

In the Internal Hellenides, the D5 structures are generally related to extensional tectonics and exhumation processes of deep structural units, either in the form of tectonic windows or metamorphic core complexes. They are recognized in the Olympos-Ossa, Rizomata and Paikon windows, as well as in the Pangaion and Cyclades metamorphic complexes (Figures 3, 4, 7, 8c, 16; [7,18,30,39,86,142,195,199,200,210,243,294]).
The D5 structures have been dated as of Oligocene-Miocene age (Figure 11a, b; e.g., [6,7,142,151]) and in the Internal Hellenides are characterized by normal detachment faults and mylonitic rocks formation in the structurally lower crustal levels. Brittle to semi-ductile deformation is recorded at the structurally higher crustal levels (e.g., the upper part of the Pelagonian nappe in northern Greece, including the Paikon window) and ductile, related to green-schist to amphibolite facies metamorphic conditions until migmatization and granite intrusions at the lower ones (e.g., the Olympos-Ossa and Rizomata windows, the Cyclades and the Rhodope Pangaion metamorphic core complexes). Here, it should be mentioned the Eocene-Oligocene space-time, where the middle Rhodope Sidironero unit was progressively exhumed during extension associated with high-temperature metamorphism, migmatization processes and granitoid intrusions (Figures 4, 17a, c; [7,17,24,40,67,197,198,199,305,306]). The cooling/exhumation path for the Sidironero unit, as it was previously referred to, was dated 42-30 Ma, the same as the lower Serbo-Macedonian Kerdylia unit (Figures 4, 17a, b; [225]). This Eocene-Oligocene extensional stage in the Rhodope province was further extended to the D5 Oligocene-Miocene extension.
The D5 stretching lineation on the S5 schistosity planes plunges in general SW- or NE-ward. Additionally, the sense of shear during the D5 event in the internal Hellenides is mainly toward SW (e.g., Pelagonian nappe and Serbo-Macedonian/Rhodope nappes pile). An opposite sense of movement toward the NE has also been observed in a few places (e.g., Olympos-Ossa window), indicating a bivergent orogenic crustal deformation during D5 (Figures 3, 4, 25; [5,6,86,142,155,156,198,210,295]). In the Cyclades massif, D5 tectonics is recorded by a main NNE-ward sense of movement related to extensional tectonics, high-temperature metamorphism and the final exhumation of the Cyclades metamorphic complex. Nevertheless, an SSW-ward sense of shear is also described for the Cycladic massif during the D5 extensional setting (Figure 19; [89,103,307]).
Contrary to the D5 structural setting in the Internal Hellenides, a fold and thrust belt, in the form of a thin-skinned type tectonics was developed in the External Hellenides during the Oligocene-Miocene, clearly related to compression and crustal stacking. Here, metamorphic processes are not observred. A main SW-ward sense of movement is also recognized by the thrust faults geometry and the folds vergence. Thrust and fold structures strike mainly NW-SE (Figures 3, 4, 24, 26). Back-thrusting top-to-the-NE is often observed in places. During the Oligocene-Miocene, the Gavrovo zone, together with the overthrust during the Eocene-Oligocene external Pindos nappe, was thrust toward SW over the Ionian zone, following the general Tertiary SW-ward orogenic migration of compression. Furthermore, the Pindos zone, possibly due to an out-of-sequence thrust fault, was thrust in places further to the west on the Ionian zone during the Early Miocene, covering completely the Gavrovo zone and its contact with the Ionian zone [25,28,29].
Additionally, HP/LT metamorphism of Oligocene-Miocene age and compressional tectonics, causing nappe stacking and crustal thickening, are recognized more south in the External Hellenides, in the Peloponnese and Crete island. (Figures 6, 19; [31,32,33,36,83,84]). Here, parts of the Ionian zone, together with the Phyllite-Quartzite unit of unknown origin, were subducted during the Oligocene-Miocene below the Gavrovo zone and metamorphosed in HP/LT conditions. So compression and extension take place again simultaneously in different geotectonic places in the Hellenides during the Oligocene-Miocene. HP/LT metamorphism and nappe stacking were progressively followed by Early-Middle Miocene ductile extensional tectonic, nappe collapse and exhumation of the high-pressure rock units in the form of a series of tectonic windows (e.g., Taygetos-Mani, Lefka Ori, Psiloritis, Dikti-Lathisi and Agios Nikolaos windows) under an isothermal decompression P/T/t-path. Sense of shear during extension and crustal exhumation was recorded top to the NNE- and SSW-ward (Figures 6, 18; [32,33,34,83,296,297,308,309,310]). At the same time, compression migrated south of the Crete island along the Hellenic active subduction zone, where the African plate is subducted under the Hellenides (Figure 19).
Figure 26. Field photos of stratigraphical sequences and tectonic structures of the External Hellenides zones: I. II. III. IV. The typical S/SW-ward asymmetrical, tight folds of the Pindos series in red Jurassic radiolarian cherts (Pindos Mt., I), intensively folded pelagic Cretaceous limestones with siliceous chert layers (Pindos Mt., II), pelagic Triassic limestones with siliceous chert layers (South Crete island, III, IV); V, VI, VII. The Early Cretaceous Pindos “first flysch”, intensively multi-folded; VIII. SW-ward Eocene-Oligocene thrust fault in the Gavrovo flysch (Pindos Mt); IX. Oligocene-Miocene out-of-sequence SW-ward thrust fault in the Pindos zone (Pindos Mt). The Triassic Pindos limestones overthrust the Jurassic Pindos radiolarian cherts, which overthrust secondary the Ionian zone during the Oligocene-Miocene, possibly due to an out-of-sequence mega-thrust fault zone; see also Figures 3, 4 (modified after [30]).
Figure 26. Field photos of stratigraphical sequences and tectonic structures of the External Hellenides zones: I. II. III. IV. The typical S/SW-ward asymmetrical, tight folds of the Pindos series in red Jurassic radiolarian cherts (Pindos Mt., I), intensively folded pelagic Cretaceous limestones with siliceous chert layers (Pindos Mt., II), pelagic Triassic limestones with siliceous chert layers (South Crete island, III, IV); V, VI, VII. The Early Cretaceous Pindos “first flysch”, intensively multi-folded; VIII. SW-ward Eocene-Oligocene thrust fault in the Gavrovo flysch (Pindos Mt); IX. Oligocene-Miocene out-of-sequence SW-ward thrust fault in the Pindos zone (Pindos Mt). The Triassic Pindos limestones overthrust the Jurassic Pindos radiolarian cherts, which overthrust secondary the Ionian zone during the Oligocene-Miocene, possibly due to an out-of-sequence mega-thrust fault zone; see also Figures 3, 4 (modified after [30]).
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4.6. Neogene-Quarternary deformational event (D6; ~20 Ma to recent)

The D6 event is mainly related to extension in all the Hellenides’ areas and took place under brittle conditions. The D6 structures overprint all the previously described structures and represent the final deformational stages of the orogen. At these stages, the compression and the orogenic evolution migrated further SW-ward now into the arcuate-form active Hellenic subduction zone south of Crete island, related to the active Hellenic volcanic arc in the central Aegean sea (Figures 19, 20; [23,188,311]).
The D6 structures are characterized by high-angle, both dip-slip and oblique-slip normal faults, as well as strike-slip faults, while some of them are related to the development of the Neogene-Quaternary basins in the Hellenides [70,293,312,313,314,315,316]. Many of the D6 faults produced significant tectonostratigraphic gaps juxtaposing higher tectonic nappes against lower ones (Figures 4, 13; [6,142,294]). Furthermore, some of the D6 faults are still active and often associated with strong earthquakes [314,317,318,319].

5. Geotectonic reconstruction of the Hellenides and Discussion

In this chapter, according to the described structural architecture, the kinematics of deformation and the main magmatic and sedimentary processes in the several Hellenides domains, we attempt to reconstruct the Alpine geotectonic evolution of the Hellenic orogenic belt (Table 1). Additionally, we discuss here the several views that dominate in the modern international bibliography about the Hellenides` geological history. We should emphasize that although much scientific researche has been published about the structural and geotectonic evolution of the Hellenides, many questions remain open. In this case, our proposal about the geotectonic evolution of the Hellenides, although based on all available geological and structural data, remains speculative in some respects.
Initially, the described ultrahigh-pressure metamorphic rocks with diamond-bearing paragenesis of ca. 200 Ma formation age [56,231] are possibly formed during the S-SW-ward dipping Paleotethys subduction zone and they trace the old suture of the Paleotethys ocean [320,321,322,323,324]. Subsdequently, during the Alpine orogeny, these ultrahigh-pressure rocks were tectonically incorporated in the upper and middle Serbo-Macedonian/Rhodope Vertiskos-Kimi and Kerdylia-Sidironero units, respectively [17,24].
Continental rifting of the Pangea super-continent and the opening of the Neotethys ocean basin (Meliata/Maliac ocean) started during the Permo-Triassic, possibly related to the closure of the Paleotethys further to the north (Figure 18; [13,20,133,146,159,277,321,322,323]. Continental rifting is associated with bimodal volcanic activity and neritic-clastic sediment deposition along the newly formed passive continental margins of Apulia and Europe, at both edges of the newly established Neotethyan ocean. The northeastern continental margin belongs to Apulia, including the Pelagonian continental domain and the southwestern continental margin belongs to the European continent, including the Vertiskos-Kimi unit of the Serbo-Macedonian/Rhodope massif. Furthermore, A-type leucocratic granitoids intruded into the Paleozoic Apulian and European basement rocks during the rifting. Carbonate neritic platform to pelagic sediments and turbidites (e.g., Dachstein, Hallstatt and Meliata Facies) were progressively deposited, from the Early Triassic to the Early-Middle Jurassic, along the continental margins of the European and Apulian plates (Figures 18, 27; [3,13,20,130,131,133,146,147,159,260,261,275,277]).
During the Early-Middle Jurassic, parts of the Triassic Meliata/Maliac Neotethys ocean, subducted towards SE in an intra-oceanic subduction regime (Figure 27a), progressed in an arcuate NW-ward convex subduction zone [11,12,13,24,43,154,164]. This geometry could be caused by a northwest-directed retreat of the subduction zone due to the roll-back of the subducting slab [11,12,24,180]. Due to this intra-oceanic subduction, which developed progressively during the Middle-Late Jurassic, in a supra-subduction position in the backarc area behind an ensimatic island arc, the Axios/Vardar ocean, part of which should also be accepted as the Paionia ocean basin (Figure 27a; [11,12,16,20,24, 142,164,175]). In the next stage, during the late Middle-Late Jurassic, the vorearc lithosphere, the island arc formation together with the Axios/Vardar ocean lithosphere and the ophiolitic mélanges (e.g., Avdella mélange) formed in deep water basins at the front of the overriding, advanced upper oceanic lithosphere, were obducted towards West onto the Apulian-Pelagonian continental margin and towards East onto the upper Serbo-Macedonian/Rhodope Vertiskos-Kimi unit, consisting a part of the European continental margin (Figure 27a, b; [6,13,17,20,24,43,154,180]). This simultaneous, opposite geometry of the emplacement kinematics on both the western and eastern sides of the Meliata-Axios/Vardar ocean basin, Apulia and Europe, respectively, was interpreted by the progressively evolved strong curvature of this Early-Middle Jurassic intra-oceanic subduction zone. (Figure 27a, b; [17,24]).
Formations of an amphibolite sole of 170-180 Ma age (Figures 23, 27a; [158,286,287]) and Middle-Late Jurassic island-arc intermediate to silicic volcanic products, tectonically incorporated in between the obducted ophiolitic belt, are related to the development of this intra-oceanic subduction framework and the formation of an ensimatic island arc (e.g., the volcanic and volcanoclastic products outcropping today in the Axios/Vardar zone and the magmatic Chortiatis series, Figures 4, 13, 14, 21, 22, 27a,b; [6,17,24,135,142,162,177]). Additionally, Late Jurassic granite intrusions in the Axios/Vardar ophiolites are interpreted as products of the magmatic activity, that took place during the progression of the Meliata/Maliac intra-oceanic subduction and the formation of the Axios/Vardar ocean (e.g., the I-type Fanos granite, Figures 3, 13, 14, 27; [164]).
The Late Jurassic calc-alkaline granites today found as orthogneisses and mylonitgneisses in the middle Rhodope Sidironero unit [201] and in the Stip-Axios massif [171,172] should also be related to the progressive stages of the supra- subduction Axios/Vardar ocean development. They represent possibly the plutonic infrastructure of the arc volcanic products (Figures 3, 4, 27a, b; [24,180]).
The Late Jurassic high-pressure metamorphism (>150 Ma), predating the D1 Late Jurassic-Early Cretaceous syn-metamorphic deformation, is ascribed to the Middle-Late Jurassic nappe stacking deformational stage. The Triassic-Jurassic sedimentary sequences of the Apulia-Pelagonia and Europe-Vertiskos/Kimi passive continental margins, together with the obducted arc-formations and ophiolitic belt, were detached outward and due to their weight, the basement rocks of both margins were buried in the deep, where they metamorphosed in high-pressure conditions (Figure 27b, c; [6,19,24,142]).
During the D1 progression, the high-pressure metamorphic basement rocks came up to shallower crustal levels and were strongly deformed simultaneously with extension and greenschist-to-amphibolite facies metamorphism, well exposed in the Pelagonian nappe as well as in the Serbo-Macedonian massif (Figures 11a, b, 17b, 27b, c; [6,13,126,127,142,149,150,151,242,275,277,298]). Furthermore, the accumulated and further strongly eroded Late Jurassic (Kimmeridgian-Tithonian) shallow-water sedimentary platform carbonates on the top of the obducted ophiolites, clearly determining the upper limit of the ophiolite emplacement in the Oxfordian-Kimmeridgian, either they emplaced on the eastern and western margins of the Pelagonian nappe or the Serbo-Macedonian Vertiskos unit, indicate a simultaneous emplacement of the ophiolites on both Apulia (Pelagonian) and European (Serbo-Macedonian) continental margins (Figures 25c, 27c; [13,143,146,148,149,150,256,257,275,277,325]). According to this evidence and taking also into account our structural works [6,142,164], as well as a lot of recent studies concerning the geodynamic evolution of Hellenides [16,17,20,24,43,154], it is inferred that all ophiolite nappes were originated from a single source and this was the Neotethyan Meliata/Maliac-Axios/Vardar ocean basin. Subsequently, the ophiolite nappes should be considered far-traveled nappes on the two different continental parts of Apulia (Pelagonian) and Europe (Serbo-Macedonian), (Figure 27b, c).
Figure 27. (Without scale). Schematic crustal-scale transects showing the structural evolution of the Hellenides during the Alpine orogeny from the Early-Middle Jurassic until the late Early Cretaceous. Has been preceded the Permo-Triassic Pangea continental rifting and the progressive Neotethyan Meliata/Maliac ocean opening, that is shown in Figure 18, 240 Ma granite ages (modified after [6,17,24,119,120,183]): a. Early-Middle Jurassic; intra-oceanic subduction in an arcuate, northwestward convex subduction zone, associated with the formation of amphibolite metamorphic sole and ophiolitic mélanges at the front of the advanced, overriding oceanic lithosphere, as well as island arc magmatism. A new oceanic lithosphere was progressively formed (the Axios/Vardar ocean) by back-arc spreading in the supra-subduction zone (SSZ). b. Middle-Late Jurassic (Oxfordian-Kimmerdgian); Arc-continent collision, ophiolite obduction, ophiolitic mélange formations at the front of the advanced ophiolites, high-pressure metamorphism, W-ward and E-ward sense of movement and crustal imbrication at the western (Pelagonian/Apulia) and eastern (Serbo-Macedonian/Europe) marginal parts of the Neotethyan realm, respectively. Subsequently, deposition of the Late Jurassic neritic sedimentary sequence on the obducted ophiolite realm. c. Late Jurassic-Early Cretaceous; Retrogression under greenschist to amphibolite facies metamorphic conditions during the Late Jurassic-Early Cretaceous and progressively possible uplifting of both continental margins (D1). Deposition of the Late Jurassic-Early Cretaceous mass-flow deposits during extension and crustal uplift. d. Early Cretaceous (D2, Barremian-Aptian); W-ward and E-ward crustal imbrication and nappe stacking at both the Pelagonian and Serbo-Macedonian continental margins, respectively. Syn-tectonic greenschist facies metamorphism. The future Late Cretaceous subduction zone along which the remnants of the Neotethyan Axios/Vardar ocean will be subducted under Europe is shown by a dotted, grey line. Abbreviations as in Figure 3.
Figure 27. (Without scale). Schematic crustal-scale transects showing the structural evolution of the Hellenides during the Alpine orogeny from the Early-Middle Jurassic until the late Early Cretaceous. Has been preceded the Permo-Triassic Pangea continental rifting and the progressive Neotethyan Meliata/Maliac ocean opening, that is shown in Figure 18, 240 Ma granite ages (modified after [6,17,24,119,120,183]): a. Early-Middle Jurassic; intra-oceanic subduction in an arcuate, northwestward convex subduction zone, associated with the formation of amphibolite metamorphic sole and ophiolitic mélanges at the front of the advanced, overriding oceanic lithosphere, as well as island arc magmatism. A new oceanic lithosphere was progressively formed (the Axios/Vardar ocean) by back-arc spreading in the supra-subduction zone (SSZ). b. Middle-Late Jurassic (Oxfordian-Kimmerdgian); Arc-continent collision, ophiolite obduction, ophiolitic mélange formations at the front of the advanced ophiolites, high-pressure metamorphism, W-ward and E-ward sense of movement and crustal imbrication at the western (Pelagonian/Apulia) and eastern (Serbo-Macedonian/Europe) marginal parts of the Neotethyan realm, respectively. Subsequently, deposition of the Late Jurassic neritic sedimentary sequence on the obducted ophiolite realm. c. Late Jurassic-Early Cretaceous; Retrogression under greenschist to amphibolite facies metamorphic conditions during the Late Jurassic-Early Cretaceous and progressively possible uplifting of both continental margins (D1). Deposition of the Late Jurassic-Early Cretaceous mass-flow deposits during extension and crustal uplift. d. Early Cretaceous (D2, Barremian-Aptian); W-ward and E-ward crustal imbrication and nappe stacking at both the Pelagonian and Serbo-Macedonian continental margins, respectively. Syn-tectonic greenschist facies metamorphism. The future Late Cretaceous subduction zone along which the remnants of the Neotethyan Axios/Vardar ocean will be subducted under Europe is shown by a dotted, grey line. Abbreviations as in Figure 3.
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During the late Early Cretaceous (D2, Barremian-Aptian; ~125-115 Ma), ongoing plate convergence and possible pull of the subducted oceanic lithospheric slab led further to outward W-SW-vergent imbrication and folding of the whole Pelagonian nappe pile, including the previously obducted ophiolites, the Late Jurassic neritic platform carbonates, the Early Cretaceous mass flow sedimentary formations and the Paleozoic basement rocks. D2 was generally related to a greenschist facies metamorphism (Figures 4, 11a, b, c, 27d; [6,43,142,149,150,154]). On the contrary, an E- to NE-ward sense of movement and imbrication has been identified for the eastern side of the Axios/Vardar ocean at the same time, as can be explained by the proposed evolution of an arcuate Jurassic subduction zone in the Neotethyan Meliata/Maliac-Axios/Vardar ocean realm (Figures 4, 17b, 27d; [17,24,180]). Anatectic melts in the Pelagonian basement during the late Early Cretaceous (~117 Ma) have also been recorded by [43,154], indicating a high-grade temperature flow in places during the D2. The Beotian flysch of Early Cretaceous age [125] and references therein), as well as some ophiolite mélange occurrences that were deposited at that time, should represent the sedimentary infill of the foreland basins formed at the front of the D2 thrust sheets. The Early Cretaceous high-pressure metamorphism (~120 Ma) referred to by [50,57] for the upper Rhodope Kimi unit could be explained as a consequence of the D2 compression and crustal thickening.
Crustal thickening due to the D2 event was followed in the Apulian/Pelagonian site by a late Early Cretaceous-Late Cretaceous extension. During the Late Cretaceous-Paleocene, the transgressive shallow water carbonate series takes place lying discordantly over all the previously described pre-Late Cretaceous sedimentary formations and structures (Figures 4, 13, 15, 28a; [6,142]). This extensional regime may have led to the continental breakup of Apulia. The break-up of Apulia was associated with the formation of a small ocean basin in the Late Cretaceous named the Pindos-Cyclades ocean, which separated the Pelagonian fragment from Apulia (Figure 28a; [22,23,24,59,117]). This new oceanic crust is supported by the dating of the protolith of ophiolites at ~80 Ma in the Cyclades area and Crete island [23,191,192].
On the contrary, in the European margin, compression and W-SW-ward thrust vergence developed during the Late Cretaceous. This compressional tectonic was related to the subduction of the Axios/Vardar ocean remnants under the European margin, including the Serbo-Macedonian/Rhodope Vertiskos-Kimi unit. Late Cretaceous subduction processes were indicated here by the existence of a Late Cretaceous magmatic arc along the European margin above the subduction zone (i.e., Strandja/Sredna-Gora massif in Bulgaria, Figures 2, 20, 28a, b; [23,24,236,266]). Furthermore, the Late Cretaceous high-to-ultra-pressure rocks (eclogites age 70-80 Ma) recognized as imbricated slices in between the Rhodope nappes strongly support this Late Cretaceous subduction process [59,117]. During the late Late Cretaceous-Paleocene structural stage, the oceanic lithosphere of the subducted Axios/Vardar ocean, including Late Jurassic to Early Cretaceous calc-alkaline arc-related granitoids and segments of European continental origin, following the subducting ocean slab, was detached E-NE-ward and accreted to the active European continental margin (Kimi unit), forming the Serbo-Macedonian/Rhodope middle Sidironero-Kerdylia unit (Figures 4, 28b).
During the Paleocene-Eocene to Early Oligocene, the D4 compressional event follows. It is related to the final stages of the Late Cretaceous subduction processes of the Axios/Vardar ocean under Europe and the subsequent development of the SW-ward vergent thrust and fold belt in the Apulian and Axios/Vardar structural units without any important metamorphic event, at least at their tectonically higher parts. On the contrary, in the Serbo-Macedonian/Rhodope province, extension, crustal uplift and exhumation processes took place simultaneously with compression and nappe stacking under ductile conditions (Figures 4, 7, 13, 14, 28b; [6,27,37,294]). During the Paleocene-Eocene, the entire Axios/Vardar ocean lithosphere was buried under the European margin and the Axios/Vardar ocean closed. As a consequence, the European plate, together with fragments of the Axios/Vardar zone units, the late Late Cretaceous high-pressure metamorphosed rocks and the Sidironero-Kerdylia unit, collided with the Apulian`s Pelagonian fragment, sliding SW-ward along segments of the Axios/Vardar oceanic remnants (Figures 4, 28b; [17,24]). Due to the collision, the European Vertiskos-Kimi unit overthrust the Triassic-Jurassic volcano-sedimentary Circum-Rhodope belt. During the D4 event, as compression and nappe stacking were advancing towards the West through time, at the Serbo-Macedanian/Rhodope nappes pile, compression and the related high-pressure metamorphism were progressively replaced by extension, nappes collapse mainly towards the SW and gradual exhumation of the higher to the deeper crustal nappes (e.g., initially with the Vertiskos-Kimi unit). The extension was now related to high-grade temperature metamorphism and migmatites creation, as well as new adakitic magmatic activity dated as of Paleocene-Eocene age (e.g., Elatia granite, Figures 4, 16, 17a, c, 28b; [51,57,197,214,219,220,224]). The high-temperature metamorphism and the magmatic activity were possibly triggered by the break-off of a deep-seated slab, maybe of the downgoing Pelagonian margin, either by delamination or convective thinning of the previously over-thickened Rhodope lithosphere, resulting in upwelling of asthenospheric material and flow heating (Figure 29b; [57,210,220]).
Figure 28. (Without scale). Schematic crustal-scale transects showing the structural evolution of the Hellenides during the Alpine orogeny from the early Late Cretaceous until the Neogene-Quaternary, CRB=Circum-Rhodope belt, 240 Ma granite ages (modified after [6,17,24,119,120,183]): e. late Early Cretaceous-Paleocene; extension during the Albian-Late Cretaceous (D3). Unconformable sedimentation of the Late Cretaceous shallow-water carbonate series terminated with the Early Paleocene internal Hellenides flysch. Progressively, the development of the Late Cretaceous subduction of the Axios/Vardar ocean remnants under the Serbo-Macedonian margin (Europe) and the formation of Late Cretaceous calc-alkaline, arc-type magmatism at the European margin, as well as the Late Cretaceous high-pressure/high-temperature metamorphic belt (HP/HT). Exhumation of the uppermost Serbo-Macedonian parts during the late Late Cretaceous-Paleocene (parts of the Vertiskos unit). Possible spreading of the ?Pindos/Cyclades small ocean basin, now between the Pelagonian and Apulia. Its future Paleocene-Eocene subduction under the Pelagonian nappe is also indicated by the grey line. f. Paleocene-Eocene/Oligocene (D4); compression, W- to SW-ward sense of movement and intense crustal imbrication of the Pelagonian nappe and Axios/Vardar zone. Final collision of Europe (Serbo-Macedonian) with the Pelagonian nappe during the Paleocene-Eocene, simultaneously with Extension and the exhumation of the unified Vertiskos-Kimi unit. Formation of the Internal Hellenides high-pressure belt of Paleocene-Eocene age (Olympos-Ossa, Cyclades) due to the subduction of the small Pindos/Cyclades ocean under the Pelagonia. Progressively, emplacement during the Eocene-Oligocene of the Pelagonian nappe (Apulia) together with the HP/LT internal Hellenides metamorphic belt on the External Hellenides Olympos-Ossa unit, which constituted the Triassic-Oligocene platform carbonate sequence of the Apulian passive margin. Simultaneously, along the Nestos ductile thrust zone, tectonic emplacement of the Sidironero-Kerdylia unit on the Pangaion unit takes place, considering the latter as equivalent to the Olympos-Ossa unit. In that case, the suture zone between the Pelagonian nappe and Olympos-Ossa unit should be the same as the tectonic contact between the Sidironero-Kerdylia and Pangaion units. So that the same suture zone continues from the Pelagonian nape until the Serbo-Macedonian/Rhodope nappe pile. Intense compression and nappe stacking in the External Hellenides Pindos and Gavrovo units. Eocene-Oligocene syn-orogenic extension in the Serbo-Macedonian/Rhodope metamorphic province associated with high-temperature metamorphism, migmatization and magmatism causes the exhumation of the Sidironero-Kerdylia unit. g. Oligocene-Miocene to Quaternary (D5, D6); HP/LT metamorphism and building of the External Hellenides Oligocene-Miocene high-pressure belt associated with compression and nappe stacking in the External Hellenides. Simultaneous syn-orogenic extension, low- to high-temperature metamorphism and magmatism in the Internal Hellenides (Olympos-Ossa, Cyclades and Serbo-Macedonian/Rhodope metamorphic province) and finale exhumation of the lowermost Rhodope Pangaion unit in the form of a metamorphic core complex, regarded as part of the External Hellenides Olympos-Ossa unit. (D6). Neogene-Quaternary: active Hellenic subduction zone and compression associated contemporaneously with extension and rapid exhumation during the Early-Middle Miocene of the external Hellenides Oligocene-Miocene HP/LT belt as a series of tectonic windows in the External Hellenides in Peloponnese and Crete island. Finally, high-angle normal and strike-slip faults, some of which are related to intramontagne basin formations and/or recent neo-tectonic activity. Abbreviations as in Figure 3.
Figure 28. (Without scale). Schematic crustal-scale transects showing the structural evolution of the Hellenides during the Alpine orogeny from the early Late Cretaceous until the Neogene-Quaternary, CRB=Circum-Rhodope belt, 240 Ma granite ages (modified after [6,17,24,119,120,183]): e. late Early Cretaceous-Paleocene; extension during the Albian-Late Cretaceous (D3). Unconformable sedimentation of the Late Cretaceous shallow-water carbonate series terminated with the Early Paleocene internal Hellenides flysch. Progressively, the development of the Late Cretaceous subduction of the Axios/Vardar ocean remnants under the Serbo-Macedonian margin (Europe) and the formation of Late Cretaceous calc-alkaline, arc-type magmatism at the European margin, as well as the Late Cretaceous high-pressure/high-temperature metamorphic belt (HP/HT). Exhumation of the uppermost Serbo-Macedonian parts during the late Late Cretaceous-Paleocene (parts of the Vertiskos unit). Possible spreading of the ?Pindos/Cyclades small ocean basin, now between the Pelagonian and Apulia. Its future Paleocene-Eocene subduction under the Pelagonian nappe is also indicated by the grey line. f. Paleocene-Eocene/Oligocene (D4); compression, W- to SW-ward sense of movement and intense crustal imbrication of the Pelagonian nappe and Axios/Vardar zone. Final collision of Europe (Serbo-Macedonian) with the Pelagonian nappe during the Paleocene-Eocene, simultaneously with Extension and the exhumation of the unified Vertiskos-Kimi unit. Formation of the Internal Hellenides high-pressure belt of Paleocene-Eocene age (Olympos-Ossa, Cyclades) due to the subduction of the small Pindos/Cyclades ocean under the Pelagonia. Progressively, emplacement during the Eocene-Oligocene of the Pelagonian nappe (Apulia) together with the HP/LT internal Hellenides metamorphic belt on the External Hellenides Olympos-Ossa unit, which constituted the Triassic-Oligocene platform carbonate sequence of the Apulian passive margin. Simultaneously, along the Nestos ductile thrust zone, tectonic emplacement of the Sidironero-Kerdylia unit on the Pangaion unit takes place, considering the latter as equivalent to the Olympos-Ossa unit. In that case, the suture zone between the Pelagonian nappe and Olympos-Ossa unit should be the same as the tectonic contact between the Sidironero-Kerdylia and Pangaion units. So that the same suture zone continues from the Pelagonian nape until the Serbo-Macedonian/Rhodope nappe pile. Intense compression and nappe stacking in the External Hellenides Pindos and Gavrovo units. Eocene-Oligocene syn-orogenic extension in the Serbo-Macedonian/Rhodope metamorphic province associated with high-temperature metamorphism, migmatization and magmatism causes the exhumation of the Sidironero-Kerdylia unit. g. Oligocene-Miocene to Quaternary (D5, D6); HP/LT metamorphism and building of the External Hellenides Oligocene-Miocene high-pressure belt associated with compression and nappe stacking in the External Hellenides. Simultaneous syn-orogenic extension, low- to high-temperature metamorphism and magmatism in the Internal Hellenides (Olympos-Ossa, Cyclades and Serbo-Macedonian/Rhodope metamorphic province) and finale exhumation of the lowermost Rhodope Pangaion unit in the form of a metamorphic core complex, regarded as part of the External Hellenides Olympos-Ossa unit. (D6). Neogene-Quaternary: active Hellenic subduction zone and compression associated contemporaneously with extension and rapid exhumation during the Early-Middle Miocene of the external Hellenides Oligocene-Miocene HP/LT belt as a series of tectonic windows in the External Hellenides in Peloponnese and Crete island. Finally, high-angle normal and strike-slip faults, some of which are related to intramontagne basin formations and/or recent neo-tectonic activity. Abbreviations as in Figure 3.
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Paleocene-Eocene extension, nappes tectonic denudation and crustal exhumation in the Serbo-Macedonian/Rhodope nappes pile take place about the same time, with compression, nappes stacking and subduction of the Cyclades/Pindos small ocean or parts of the Pindos deep sea basin under the Pelagonian fragment. The same subduction event in the Olympos-Ossa area further to the North is described as an A-type subduction (Figures 4, 7, 19, 28a, b; [24,47,86,93,155]). Simultaneously, further to the west, in the External Hellenides, the external parts of the Pindos zone were strongly SW-ward imbricated and folded. Thin-skinned tectonics characterized this compressional tectonic of the External Hellenides, building up the structurally complicated Pindos nappe (Figures 4, 26, 27b). The Internal Hellenides, Paleocene-Eocene Ampelakia-Cyclades high-pressure belt was formed during this subduction processes (Figures 2, 7, 28a, b; [47,48,86,93]). [114], [17], [24] suggest the continuation of this Paleocene-Eocene high-pressure belt until the Rhodope Sidironero unit, additionally interpreting the lowermost Rhodope Pangaion unit as an exhumed part of the External Hellenides, equivalent to the Olympos-Ossa carbonate platform, which covers the Apulian basement. Paleocene-Eocene age`s (ca. 50-42Ma) high- to ultrahigh-pressure rocks were recognized imbricated in the Sidironero unit [51,53,57] and in the Nestos ductile thrust zone, along which the Sidironero unit overthrusts the Pangaion unit [17,24,117,223]. The same tectonostratigraphy is also found southwestern-southern in the Olympos-Cyclades nappes stack, where the Paleocene-Eocene Ampelakia-Cyclades high-pressure belt with the Pelagonian fragment overthrusts the External Hellenides Olympos-Ossa and Cyclades units [39,47,48,86]. As a result, the Ampelakia/Olympos-Cyclades thrust zone should be the same, with the Nestos shear zone continuing under the Pelagonian and Serbo-Macedonian/Rhodope nappe stack, as was also proposed by [17,24,114]. In this scenario, it should be considered that the lower Rhodope Pangaion unit is really equivalent to the Olympos-Ossa unit and the Apulia plate continues below the Serbo-Macedonian/Rhodope nappes stack (Figure 28).
Further Eocene-Oligocene nappe stacking and crustal thickening to the West, in the External Hellenides, are associated with the final emplacement of the Ampelakia-Cyclades high-pressure rocks and the Pelagonian nappes stack on the External Hellenides Gavrovo carbonate platform. Compression again took place with simultaneous extension at the structurally internal higher Hellenides nappes and exhumation now of the Sidironero-Kerdylia unit (Figures 4, 17a, 26, 27b; [17,24,40,143,198,199,295]). At the same time, new magmatic activity took place in the whole Serbo-Macedonia/Rhodope province (e.g., Vrondou and Xanthi granites, volcanic rocks in between the turbidites deposits of the Thrace basin, etc.; Figures 3, 4, 16; [7,114,197,207,210,214,219,220,326]). Eocene syn-extensional high-temperature metamorphism and migmatization processes in the middle Serbo-Macedonia/Rhodope Sidironero-Kerdylia unit, ascribed to mantel delamination processes (Figures 17a, c, 28; [17,24,51,198,219,220]). The higher-grade metamorphosed Rhodope units during the Tertiary, compared with the Pelagonian Apulian`s fragment, associated with migmatization processes, granitoid intrusions and deep crustal exhumation, could be well explained due to their deeper tectonic position to the East in the Hellenides geotectonic frame.
Here, we should also mention that the Kesebir/Kardamos and Kechros domes in the Rhodope province, which have been placed on many geological maps as equivalent to the lower Rhodope Pangaion unit, do not show clear structural and compositional relationships with the Pangaion unit. They do not contain the thick carbonate sequence as the Pangaion unit; they are overlain directly by the upper Rhodope Kimi unit, equivalent to the Serbo-Macedonian Vertiskos unit and the most important difference to the Pangaion unit is that their cooling/exhumation path was dated at the Eocene-Oligocene (~40-30 Ma; 225), which is the same cooling/exhumation age given for the middle Rhodope Sidironero unit [17,24,40,225]. Moreover, apatite fission-track analyses (AFT) for the Sidironero unit and the Kesebir/Kardamos and Kechros domes gave about the same ages, ~26 Ma and ~30 Ma, respectively. Apatite fission-track ages for the Rhodope lower Pangaion dome range between 15 and 10Ma, which are too younger than the Kesebir/Kardamos and Kechros domes (Figure 4; [225]). Furthermore, in the tectonic contact between the upper Rhodope Kimi unit and these two domes are tectonically the Late Cretaceous high-pressure rocks; a tectonic contact that has been described as the tectonic contact between the upper Kimi and middle Sidironero Rhodope units. Regarding the described structural and compositional features of both metamorphic domes, the Kesebir/Kardamos and Kechros domes fit more to be equivalents to the Sidironero-Kerdylia unit, as illustrated on our maps and cross-sections (Figures 3, 4).
During the Oligocene-Miocene (D5 event), when Europe had already collided with Apulia and Pelagonia, extension under brittle conditions took place in the structurally higher Internal Hellenides nappes levels and ductile deformation dominated in the structurally deeper levels. Ductile deformation was associated with mylonites formation, low- to high-temperature metamorphism and new magmatic activity and migmatization. So that in detail, brittle conditions dominate in the upper parts of the Pelagonian nappe pile, while ductile conditions are recognized in the lower-most parts of the Pelagonian nappe, in the lower Rhodope Pangaion unit and the Paleocene-Eocene high-pressure Ampelakia-Cyclades metamorphic belt (Figures 7a, b, 11a, b, 17a, b; [5,6,7,86,155,156,294]). The former over thickened lithospheric crust collapsed along ductile shear zones and normal detachment faults with a main top-to-the-SW sense of movement (e.g., Olympos-Ossa and Strymon-valley detachment faults; Figures 3, 4, 19, 28b, c; [5,48,86,156,207,210,295]) or with a main top-to-the-N-NE sense of movement (e.g., Cyclades detachment faults; Figures 9, 19; [10,89,188,327]). Nevertheless, an opposite sense of movement has also been described in some cases, e.g., in the Olympos-Ossa province top-to-the-NE and the Cyclades area top-to-the-S-SW, indicating here a bivergent geometry of the extensional tectonics and a bulk coaxial type of deformation. This collapse caused the formation of several tectonic windows and/or metamorphic core complexes, where the upper structural series of the Apulia plate were exhumed under the Pelagonia and the Internal Hellenides nappes stack (e.g., Olympos, Rizomata and Paikon windows, Pangaion and Cyclades metamorphic core complexes, Figures 3, 4, 7, 16, 28b, c; [6,7,30,38,47,86,89,95,142,155,205,210,295,300,328,329,330]). In the Olympos-Ossa area, the exhumation was very rapid and took place under an isothermal decompression path, while in the Rhodope and Cyclades provinces, the exhumation followed a decompression path with an initially increasing temperature gradient (Figure 8a, b; [6,38,47,48,51,88,89,95,207,210]). According to our recent works, a constrictional type deformation (i.e., NW-SE compression along the Y-axis of the strain ellipsoid, Y<1) characterizes in places the Tertiary tectonic activity in the Internal Hellenides nappes. Flattening type of deformation (Y>1) was also calculated in other places, highlighting this complicated architecture and evolution of the deformation [7,30,156,195,197,198]. Finally, according to our descriptions concerning the structural and geotectonic evolution of the Olympos-Ossa and Rizomata windows, the lowermost Rhodope Pangaion and Cyclades metamorphic core complexes, they are joined with the same tectonostratigraphy and composition of geological units as well as structural evolution.
Simultaneously, with the Oligocene-Miocene extension, nappes collapse and exhumation processes in the Olympos-Ossa-Cyclades and Serbo-Macedonian/Rhodope provinces, new compression and SW-wards verging nappe stacking developed at the External Hellenides (Gavrovo and Ionian zones) associated with new subduction processes and the formation of the External Oligocene-Miocene high-pressure belt, visible in Peloponnese and Crete island (Figures 3, 4, 6, 26, 27c). In Peloponnese and Crete island Oligocene-Miocene nappe stacking was progressively followed again by new Early-Middle Miocene bulk coaxial extension and nappes denudation along ductile shear zones and normal detachments faults, causing a series of Miocene tectonic windows and metamorphic core complexes, where the Oligocene-Miocene high-pressure rocks of the External Hellenides, represented by the Plattenkalk series (? Ionian zone) and Phyllite-Quartzite series (of unknown origin), were exhumed from under the whole Hellenides nappes stack and an isothermal decompression P-T metamorphic path (e.g., Taygetos window, Leyka Ori window, Psiloritis window, Figures 2, 3, 4, 28c; [31,32,33,34,83,296,297,328,331,332,333,334]).
During the Neogene-Quaternary, a brittle extensional deformation dominated the entire Hellenides belt. It was associated with high-angle normal and oblique strike-slip faults (D6 event), in some cases expressed by transpressional or transtensional tectonics (Figures 3, 4, 12, 14, 28c; [7,259,314]), dismembering all pre-Neogene tectonic units and structures. Now, in the present day, the convergence and compression migrated toward the SW, being directed in the active Hellenic subduction zone, where the African plate is subducting under the Hellenides towards the NE, forming the active Hellenic subduction zone and the active volcanic arc of the Aegean sea (Figures 2, 4, 20, 28c).
However, we should also refer to the evolution of the two syn- to late-orogenic sedimentary basins of the Meso-Hellenic trough and the Thrace basin. We consider these very important for the final reconstruction of the Alpine Hellenides orogenic belt. They are mentioned as molassic-type basins, filled with turbiditic and clastic sediments that lived through the Tertiary deformation history of the Hellenides. The Meso-Hellenic trough was defined as a polyhistory strike-slip and piggy-back basin behind the Tertiary External Hellenides thrust and fold belt. The Thrace basin was described as a supra-detachment basin in the Rhodope province. The deposition history of the Meso-Hellenic trough starts during the Early-Middle Eocene and ends at the Early-Middle Eocene. Respectively, the Thrace basin is filled by Eocene-Oligocene turbidites intercalated with acid to intermediate volcanic products of the same age. Younger Miocene basic magmatic products also intrude into the Paleogene deposits of the basin (Figures 1, 2, 3, 4; [60,61,62,63,64,65,66,67,243]).
The described orogenic outward Tertiary W- to SW-ward migration of the dynamic peer compression vs. extension, related respectively to nappes stacking vs. crustal uplift or to contemporaneous action of compressional and extensional tectonics in the several parts of the Hellenic orogenic belt (e.g., External and Internal Hellenides), could be well explained by a retreating subduction zone and roll-back of the subducted Pindos-Cyclades lithospheric slab and also lithospheric mantel delamination [4,6,9,18,24,188,218,219,265,335].

6. Conclusions

I. 
The Alpine structural evolution of the Hellenides starts during the Permo-Triassic with the continental rifting of the Pangea super-continent and the progressive opening of the Neotethyan Meliata/Maliac-Axios/Vardar ocean realm
II. 
Alpine deformation and metamorphism are recorded in a multiphase deformational setting from the Early-Middle Jurassic to the present day. Six main (D1 to D6) deformational events were identified that followed, at their early stages, intra-oceanic subduction and ophiolite obduction. Compression, nappe stacking, crustal thickening and high-pressure metamorphism alternated progressively through time with extension, orogenic collapse, mylonites formation and low- to high-temperature metamorphism associated with migmatites formation and granite intrusions in places. Furthermore, the extension is leading to crustal thinning, uplift and exhumation of deep crustal levels as tectonic windows or metamorphic core complexes (e.g., Olympos-Ossa window, Cyclades and Rhodope Pangaion metamorphic core complexes, Cretan and Peloponnese windows). An S-to SW-ward migration of the dynamic peer compression vs. extension is clearly recognized during the Tertiary in the Hellenides. In any case, extension and crustal uplift follow compression and nappe stacking or they take place simultaneously but in different places of the orogen. Today, compression migrated along the Hellenic active subduction zone south of the Crete island.
III. 
The ophiolite belts, including the Mid-Late Jurassic ophiolitic mélanges, in the Hellenides are considered far-traveled nappes that originated from a single source ocean, which was the Neotethyan Meliata/Maliac-Axios/Vardar ocean basin. Nappe thrusting and ophiolite overriding were started during the Early-Middle Jurassic, related to the evolution of an intra-oceanic E-ward dipping subduction in the Meliata/Maliac ocean. The upper limit of the ophiolite emplacement on the Apulian (Pelagonia) and European (Vertiskos-Kimi unit) continental margins, respectively, is the Callovian-Oxfordian. The Axios/Vardar ocean closed finally during the Paleocene-Eocene, subducting totally under the European continental margin. As a result, the Vertisko-Kimi unit, as part of the European margin, overthrusts during the late Late Cretaceous-Paleocene the Sidironero-Kerdylia unit, composed of imbricated continental parts and pieces of the Axios/Vardar NE-ward subducted lithosphere. Subsequently, Europe collided with the Pelagonian fragment of Apulian origin.
IV. 
The Axios/Vardar zone is allochthonous, intensively SW-ward imbricated by the Paleocene-Eocene compressional step-stage. The Axios/Vardar ophiolites along the Axios/Vardar zone form the exposed surface parts of the deep-seated Axios/Vardar suture, which is traced at depth in between the Rhodopes nappes. The Internal Hellenides Serbo-Macedonian/Rhodope thrust stack is rooted along this Axios/Vardar suture.
V. 
The lower-most Pangaion Rhodope unit should be the Apulian plate`s eastern marginal part. It was underthrusted along the unified Nestos-Ampelakia-Cyclades ductile thrust zone during the Eocene-Oligocene below the Pelagonian and the Serbo-Macedonian/Rhodope nappes stack, following the Paleocene-Eocene subduction of the small Pindos-Cyclades ocean under the Pelagonian fragment. This model, in agreement with Jahn-Awe et al. 2010 and Froitzheim et al. 2014 assumes an E-NE-ward, far-traveled underthrusting of the Apulia plate below the Pelagonian and the Serbo-Macedonian/Rhodope nappes pile. Finally, the lower-most Pangaion Rhodope unit exhumed as a metamorphic core complex below the Serbo-Macedonian/Rhodope nappes stack during the Oligocene-Miocene extensional tectonics of the Internal Hellenides and the broader Aegean region. Respectively, the Serbo-Macedonian/Rhodope nappes were exhumed progressively under an extensional regime, initially with the exhumation of the Vertiskos-Kimi unit during the Paleocene-Eocene, followed by the exhumation of the Sidironero-Kerdylia unit during the Eocene-Oligocene. On the contrary, during the Tertiary in the External Hellenides, compression and nappe thrusting dominate, migrating S-SW-ward until the today active Hellenic subduction zone, causing also the Oligocene-Miocene subduction processes. Moreover, Oligocene-Miocene HP/LT metamorphism is recognized between the External Hellenides in the southern Peloponnese and Crete island (Plattenkalk series and Phyllit-Quartzite unit). Here, the Early-Middle Miocene extension followed, causing a series of tectonic windows and metamorphic core complexes.
VI. 
During the Eocene-Oligocene the Pelagonian nappe with the obducted ophiolite nappes and the Ampelakia-Cyclades Paleocene-Eocene high-pressure belt were emplaced on the External Hellenides Gavrovo carbonate platform (Apulia plate). The latter exhumed subsequently during the Oligocene-Miocene extension, forming the Olympos-Ossa tectonic window as well as the Cyclades and Pangaion metamorphic core complexes simultaneously with the Oligocene-Miocene compression, subduction and SW-S-ward nappes stacking in the External Hallenides.
VII. 
A retreating subduction zone and rollback of the subducted Pindos-Cyclades lithospheric slab and/or mantel delamination processes, could well explain the Tertiary S- to SW-ward migration of the compressional and extensional tectonics. Compression is related to nappes stack and crustal thickening, extension to nappes denudation, crustal uplift, and crustal thining. Extension normally follows compression.

Funding

This research received no external funding.

Data Availability Statement

Not applicable.

Acknowledgments

Dedicated to my students of the School of Geology of the University of Thessaloniki GR, as well as to my teachers, friends and colleagues with whom I collaborated a long time of about 50 years. I can` t forget the nice field work days and the endless discussions about the geological problems and orogenic processes.
Extension vs. Compression

Conflicts of Interest

The authors declare no conflict of interest.

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Table 1. Summarized the Alpine structural evolution of the Hellenides.
Table 1. Summarized the Alpine structural evolution of the Hellenides.
1. Permo-Triassic: continental rifting; bimodal volcanism and A-type granite intrusions; starts the opening of the Neotethys ocean
2. Triassic-Jurassic: passive margins; extension and sedimentation
3. Early-Middle Jurassic: intra-oceanic subduction; amphibolite metamorphic sole; island arc magmatism during the Middle Jurassic; Middle-late Middle Jurassic development of the Axios/Vardar ocean in a supra-subduction zone
4. Middle-Late Jurassic: W-SW- and E-NE-ward ophiolite obduction and imbrication regime; high-pressure metamorphism in the two continental margins, Apulian`s and European`s, at the western and eastern edges of the Neotethian ocean, respectively; deposition of ophiolitic tectono-sedimentary melanges in basins at the front of the advanced ophiolitic belts
5. Late Jurassic: deposition of neritic platform carbonate on top of the obducted ophiolites
6. Late Jurassic-Early Cretaceous: greenschist to amphibolite facies metamorphism, possibly during extension and crustal uplift (D1); deposition during the Early Cretaceous of neritic and mass flow clastic sediments from the eroded Late Jurassic platform carbonates
7. Late Early Cretaceous (Barremian-Aptian): compression; mainly W- and E-ward imbrication and folding; syn-tectonic greenschist facies metamorphism (D2)
8. late Early Cretaceous-Late Cretaceous to Paleocene: Compression stopped; extension late Early Cretaceous-Late Cretaceous (D3); possible formation of a small ocean basin western of the Pelagonia, the Pindos-Cyclades ocean basin; Late Creaceous carbonate transgression terminated with the Paleocene Internal Hellenides flysch; subduction of the Axios/Vardar ocean remnants under the European margin and Late Cretaceous high-pressure metamorphism and compression; the middle unified Serbo-Macedonian/Rhodope Sidironero-Kerdylia unit was underthrust below the Vertiskos-Kimi unit;
9. Paleocene to Early Oligocene (D4): compression; mainly W-SW ward sense of movement; final closure of the Axios/Vardar ocean and collision of Europe with Apulia/Pelagonia during the Paleocene-Eocene; imbrication and nappes stack; strong imbrication of the Axios/Vardar units; HP metamorphism of the Internal Hellenides high-pressure belt (Ampelakia and Cyclades units) during the Paleocene-Eocene during subduction processes under the Pelagonia; progressively, emplacement of the Pelagonian and Sidironero-Kerdylia nappes together with the HP metamorphic belt on the External Hellenides during the Eocene-Oligocene; synorogenic extension in the Serbo-Macedonian/Rhodope metamorphic province associated with high-temperature metamorphism, migmatisation, granitoid intrusions and progressively crustal exhumation, initially of the Vertiskos-Kimi unit (Paleocene-Eocene) and then of the Kerdylia/Sidironero unit (Eocene-Oligocene)
10. Oligocene-Miocene (D5): HP/LT metamorphism in the External Hellenides belt associated with compression and nappe stacking in the External Hellenides, Crete island and Peloponnese; syn-orogenic extension associated with low-grade metamorphism in the Olympos-Ossa area and high- to medium-grade metamorphism, migmatization and granitoid intrusions in the Cyclades and Serbo-Macedonian/Rhodope metamorphic provinces; crustal exhumation of the Olympos-Ossa and Rhodope Pangaion units
11. Neogene-Quaternary: compression along the active Hellenic subduction zone, south of Crete island; ductile extension and exhumation of the Peloponnese and Cretan tectonic windows; brittle extension and intramontagne basin formation in the mainland (D6); Recent neo-tectonic activity
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